Bjarte Jordal





Focusing on integrative taxonomy, phylogeny and evolution of bark and ambrosia beetles and their origin within the greater scheme of weevils. These beetles show interesting variation in feeding preferences and in reproductive biology. Bark and ambrosia beetles are therefore ideal for comparative analyses, tracing the origin of key evolutionary events related to host plant associations and to changes in reproductive modes. My research focuses on two main areas:

  • Taxonomy: extensive field work in various parts of the tropics results in the discovery of new species. A particular focus is directed on the Afrotropical fauna with the goal to revise all genera and more than 1500 species. Additional research includes the Neotropical genus Scolytodes. Taxonomic work is integrative, relying to some extent on molecular markers, enabling test of biogeographical hypotheses.
  • Phylogenetics: the exact position of Scolytinae and Platypodinae within the wider scheme of weevils has turned out to be a major challenge in weevil classification. With optimization of 20 nuclear markers over the past years the resolution is still insufficient; we will continue to search for a predictive phylogeny using AHE (anchored hybrid enriched) data.

BIO233 Insect diversity and biology

BIO301 Topics in biodiversity, ecology and evolution


  1. Jordal BH. 2024. An integrated taxonomic revision of Ctonoxylon (Coleoptera, Curculionidae, Scolytinae) reveals new Malagasy species from multiple recent colonizations of the island. ZooKeys, 1203: 95–130.
  2. Jordal BH, Smith SM. 2024. The first record of the Neotropical Cnemonyx minor (Curculionidae, Scolytinae) in Africa. Entomologist’s Monthly Magazine, in press.
  3. Jordal BH. 2024. Integrated taxonomy, biology and biogeography of the Afrotropical genus Xyloctonus  (Coleoptera, Curculionidae, Scolytinae). Deutsche Entomologische Zeitschrift, 71: 67-84.
  4. Senickzak S, Senickzak A, Jordal BH. 2024. Morphological ontogeny, ecology and biogeography of Fuscozetes fuscipes (Acari, Oribatida, Ceratozetidae). Animals, 14: 538;
  5. Bickerstaff JRM, Jordal BH, Riegler M. 2023. Two sympatric lineages of Australian Cnestus solidus share Ambrosiella symbionts but not Wolbachia. Heredity,
  6. Jordal BH. 2023. An illustrated key and new records of the Afrotropical bark beetle genus Cryphalomimus Eggers (Coleoptera, Curculionidae). Zootaxa, 5353 (6): 595–600. DOI: 10.11646/zootaxa.5353.6.6
  7. Jordal, B.H. 2023. Glostatina, a new xyloctonine subtribe for Glostatus (Coleoptera: Curculionidae), based on clear genetic and morphological differences. European Journal of Entomology, 120, 199-232. DOI: 10.14411/eje.2023.025
  8. Jordal BH. 2022. Sexual dimorphism and phylogenetic position of Chilodendron (Coleoptera, Scolytinae) – a long isolated lineage endemic to Madagascar. European Journal of Entomology, 119: 152–158. DOI: 10.14411/eje.2022.017
  9. Jordal BH. 2022. The strongly dimorphic bark beetle genus Pseudomicracis (Coleoptera, Scolytinae) in Madagascar – an integrated taxonomic revision. Zootaxa, 5125 (3): 325–343.
  10. Jordal BH. 2021. New species and records of Pseudolanurgus (Coleoptera: Curculionidae, Scolytinae) from Tanzania and Madagascar. Zootaxa, 5072 (6): 592–598.
  11. Seniczak A, Seniczak S, Graczyk R, Kaczmarek S, Jordal BH, Kowalski J, Djursvoll P, Roth S, Bolger T. 2021. A Forest Pool as a Habitat Island for Mites in a Limestone Forest in Southern Norway. Diversity, 13, 578.
  12. Jordal BH. 2021. An integrated taxonomic revision of Diplotrichus (Coleoptera, Scolytinae) supports a Malagasy origin and single colonization of South Africa. Zootaxa, 5047 (2): 101–122.
  13. Seniczak A, Seniczak S, Starýc J, Kaczmarek S, Jordal BH, Kowalski J, Roth S, Djursvoll P, & Bolger T. 2021. High diversity of mites (Acari: Oribatida, Mesostigmata) supports high conservation value of a broadleaf forest in eastern Norway. Forests, 12: 1098.
  14. Jordal BH. 2021. The mainly South African genus Lanurgus revised (Coleoptera, Scolytinae). Zootaxa, 5027(1): 087–106.
  15. Jordal BH. 2021. Small, uniform, and rarely collected – an integrated taxonomic revision of Afromicracis bark beetles (Coleoptera, Scolytinae). Zootaxa, 4981(1): 070-088.
  16. Peris, D, Delclòs X. & Jordal BH. 2021. Origin and evolution of fungus-farming in wood-boring Coleoptera – a paleontological perspective. Biological Reviews,
  17. Seniczak A, Niedbała W, Iturrondobeitia JC, Seniczak S, Roth S & Jordal BH. 2021. Type of broadleaf forest matters most for ptyctimous mite communities (Acari, Oribatida) in Norway. Biodiversity and Conservation,
  18. Eliassen JM & Jordal BH. 2021. Integrated taxonomic revision of Afrotropical Xyleborinus reveals a recent species-rich radiation in Madagascar. Insect Systematics and Diversity, 5 (3): 4.
  19. Jordal BH. 2021. Revision of Dendrochilus (Coleoptera, Scolytinae) – with description of two new species from Tanzania. Zootaxa, 4969: 587–593.
  20. Jordal BH. 2021. Laximicracis — a new genus of Afrotropical Micracidini beetles (Coleoptera, Scolytinae), Zootaxa, 4966: 91–96.
  21. Jordal BH. 2021. Molecular and morphological revision of Afrotropical Hypoborini (Coleoptera, Scolytinae) reveal novel bark beetle taxa and narrow geographical endemism. European Journal of Entomology, 118: 90-110.
  22. Cognato AI, Smith SM & Jordal BH. 2021. Patterns of host tree use within a lineage of saproxlic snout-less weevils (Coleoptera: Curculionidae: Scolytinae: Scolytini). Molecular Phylogenetics and Evolution, Feb 17:107107. doi: 10.1016/j.ympev.2021.107107. Epub ahead of print. PMID: 33609714.
  23. Jordal BH. 2021. A phylogenetic and taxonomic assessment of Afrotropical Micracidini (Coleoptera, Scolytinae) reveals a strong diversifying role for Madagascar. Organisms Diversity and Evolution, 21: 245–278.
  24. Peris D, Janssen K, Barthel HJ, Bierbaum G, Delclòs X, Peñalver E, Solórzano-Kraemer MM, Jordal BH & Rust J. 2020. DNA from resin-entombed organisms: past, present and future. PLoS One, 15(9): e0239521;
  25. Jordal BH & Smith SM 2020. Scolytodes from Ecuador:  40 new species, and a molecular phylogenetic guide to infer species validity. Zootaxa, 4813: 1–67.
  26. Johnson AJ, Hulcr J, Knížek M, Atkinson TH, Mandelshtam M, Smith SM, Cognato AI, Park S, You L & Jordal BH. 2020. Revision of the bark beetle genera within the former Cryphalini (Curculionidae: Scolytinae). Insect Systematic and Diversity, 4(3):  1-81. https://doi: 10.1093/isd/ixaa002
  27. Jordal BH & Tischer M. 2020. Genetics of the widespread Afrotropical ambrosia beetle complex Xyleborus principalis reveals morphological polymorphism and new synonymies. International Journal of Tropical Insect Science, 40: 707–715.
  28. Cognato AI, Sari G, Smith SM, Beaver RA, Li Y, Hulcr J, Jordal BH, Kajimura H, Lin C-S, Pham T, Singh S & Sittichaya W. 2020. Multiple DNA loci for the identification of Southeast Asian ambrosia beetle species (Curculionidae: Scolytinae: Xyleborini). Frontiers in Ecology and Evolution, 8: 27.
  29. Mayers CG, Harrington TC, Masuya H, Mcnew DL, Jordal BH, Shih H-H, Roets F, Kietzka GJ. 2020. Patterns of coevolution of ambrosia beetle mycangia and the Ceratocystidaceae, with five new fungal genera and seven new species. Persoonia, 44: 41–66.
  30. Seniczak A, Seniczak S & Jordal BH. 2019. Molecular and ontogeny studies clarify systematic status of Chamobates borealis (Acari, Oribatida, Chamobatidae): an integrated taxonomy approach. Systematic and Applied Acarology, 24: 2409–2426.
  31. Seniczak A, Bolger T, Roth S, Seniczak S, Djursvoll P, Jordal BH. 2019. Diverse mite communities (Acari) from a rich broadleaf forest in Western Norway. Annales Zoologici Fennica, 56: 121–136.
  32. Jordal BH & Kirkendall LR. 2019. Rainforest and cloud forest Scolytodes (Curculionidae, Scolytinae, Hexacolini) from the ALAS inventory in Costa Rica: new species, new synonymy, new records. ZooKeys 863: 1-34.
  33. Jordal BH. 2019. The bark and ambrosia beetle (Coleoptera, Scolytinae) collection at the University Museum of Bergen – with notes on extended distributions in Norway. Norwegian Journal of Entomology, 66: 19–31.
  34. Mally R, Hayden JE, Neinhuis C, Jordal BH & Nuss ME. 2019. The phylogenetic systematics of Spilomelinae and Pyraustinae (Lepidoptera: Pyraloidea: Crambidae) inferred from DNA and morphology. Arthropod Systematics & Phylogeny, 77: 141–204.
  35. Jordal BH. 2018. Hidden gems in museum cabinets and drawers:  new species and new distributional records of Scolytodes (Coleoptera: Scolytinae). Zootaxa, 4504: 76-104.
  36. Cognato AI, Jordal BH & Rubinoff D. 2018. Ancient wander-lust leads to diversification of endemic Hawaiian Xyleborus species. Insect Systematics and Diversity, 2: 1-9.
  37. Johnson AJ, McKenna DD, Jordal BH, Cognato AI, Smith-Cognato SM, Lemmon AR, Lemmon EL & Hulcr J. 2018. Phylogenomics clarifies repeated origins of evolutionary innovations in bark beetles. Molecular Phylogenetics and Evolution, 127: 229–238.
  38. Mugu S, Pistone D & Jordal BH. 2018. New molecular markers resolve the phylogenetic position of the enigmatic wood-boring weevils Platypodinae (Coleoptera: Curculionidae). Arthropod Systematics & Phylogeny, 76: 45-58.
  39. Jordal BH. 2018. The smallest known species of Afrotropical Scolytoplatypus – with unique features and isolated phylogenetic position. ZooKeys, 749: 125-130.
  40. Pistone D, Gohli J & Jordal BH. 2018. Molecular phylogeny of bark and ambrosia beetles (Curculionidae: Scolytinae) based on 18 molecular markers. Systematic Entomology, 43: 387-406.
  41. Storer C, Payton A, McDaniel S, Jordal BH, Hulcr J. 2017. Cryptic genetic variation in an inbreeding and cosmopolitan pest, Xylosandrus crassiusculus, revealed using ddRADseq. Ecology and Evolution, 7:10974–10986. DOI: 10.1002/ece3.3625.
  42. Jordal BH. 2017. Ancient diversity of Afrotropical Microborus:  three endemic species – not one widespread. ZooKeys, 710 33-42.
  43. Johnson, Knížek M, Atkinson TH, Jordal BH, Ploetz R & Hulcr J. 2017. Resolution of a global mango and fig pest identity crisis. Insect Systematics and Diversity, 1: ixx010.
  44. Kambestad M, Knutsen IL, Kirkendall LR & Jordal BH. 2017. Cryptic and pseudo-cryptic diversity in the world’s most common bark beetle – Hypothenemus eruditus. Organisms Diversity and Evolution, 17: 633-652.
  45. Gohli J, Kirkendall LR, Smith S, Cognato AI, Hulcr J & Jordal BH. 2017. Biological factors contributing to bark and ambrosia beetle species diversification. Evolution, 71: 1258-1272. DOI: 10.1111/evo.13219.
  46. Gohli J & Jordal BH. 2017. Explaining biogeographic range size and measuring its effect on species diversification in bark beetles. Journal of Biogeography, 44: 2132–2144.
  47. Stouthamer R, Rugman-Jones P, Thu PQ + 17 authors. 2017. Tracing the origin of a cryptic invader: phylogeography of the Euwallacea fornicatus (Coleoptera: Curculionidae: Scolytinae) species complex. Agricultural and Forest Entomology, 19: 366-375. DOI: 10.1111/afe.12215.
  48. Jordal BH & Kaidel J. 2017. Phylogenetic analysis of Micracidini demonstrates a single Trans-Atlantic disjunction and inclusion of Cactopinus in the American clade. The Canadian Entomologist, 149: 8-25.
  49. Gohli J, Selvarajah TS, Kirkendall LR & Jordal BH. 2016. Globally distributed Xyleborus species reveal recurrent intercontinental dispersal in a landscape of ancient worldwide distributions. BMC Evolutionary Biology, 16(1): 37.
  50. Pistone D, Mugu S & Jordal BH. 2016. Genomic mining of phylogenetically informative novel nuclear markers in bark and ambrosia beetles (Curculionidae. PLoS One, Sep 26; 11(9): e0163529. doi: 10.1371/journal.pone.0163529.
  51. Jordal BH. 2016.The Book of Beetles – A Life-Size Guide to Six Hundred of Nature’s Gems, edited by Patrice Bouchard. The Quarterly Review of Biology, 91: 104-105.
  52. Jordal BH. 2015. Molecular phylogeny and biogeography of the weevil subfamily Platypodinae reveals evolutionarily conserved range patterns. Molecular Phylogenetics and Evolution, 92: 294-307.
  53. Jordal BH. 2015. Insekter sprer seg nordover. Årbok for Universitetsmuseet i Bergen 20: 96-97.
  54. Kirkendall LR, Biedermann PHW & Jordal BH. 2015. Diversity and evolution of Bark Beetles. In: Bark Beetles: Biology and Ecology of Native and Invasive Species (Vega & Hofstetter, eds).
  55. Hulcr J, Atkinson TH, Cognato AI, Jordal BH & McKenna DD. 2015. Morphology, Taxonomy and Phylogenetics of Bark Beetles. In: Bark Beetles: Biology and Ecology of Native and Invasive Species (Vega & Hofstetter, eds).
  56. Kostovcik M, Bateman C, Kolarik M, Stelinski L, Jordal BH & Hulcr J. 2015. The ambrosia symbiosis is specific in some species and promiscuous in others: evidence from high-throughput community sequencing. The ISME Journal, 9 (1): 126-138.
  57. Jordal BH. 2015. Beetles of Eastern North America, by Arthur Evans. The Quarterly Review of Biology, 90 (2): 233-234.
  58. Jordal BH, Smith SM & Cognato AI. 2014. Weevil classification as a data-driven science: leaving opinion behind. ZooKeys, 439: 1-18.
  59. Gillett CPDT, Crampton-Platt A, Timmermans MJTN, Jordal BH, Emerson BC & Vogler AP. 2014. Bulk de novo mitogenome assembly from pooled total DNA elucidates the phylogeny of weevils (Coleoptera: Curculionoidea). Molecular Biology and Evolution, 31 (8): 2223-2237.
  60. Jordal BH & Kambestad M.2014. DNA barcoding of bark and ambrosia beetles reveals excessive NUMTs and consistent east-west divergence across the Palearctic. Molecular Ecology Resources, 14: 7-14.
  61. Jordal BH. 2014. Cossoninae. In: Handbook of Zoology, Band IV Arthropoda: Insecta. Part 38: Coleoptera, Beetles, Vol. 3 (eds. Leschen RAB, Beutel R), pp 345-349. deGruyter Press.
  62. Jordal BH. 2014. Scolytinae. In: Handbook of Zoology, Band IV  Arthropoda: Insecta. Part 38: Coleoptera, Beetles, Vol. 3 (eds. Leschen RAB, Beutel R), pp 349-358. deGruyter Press.
  63. Jordal BH. 2014. Platypodinae. In: Handbook of Zoology, Band IV  Arthropoda: Insecta. Part 38: Coleoptera, Beetles, Vol. 3 (eds. Leschen RAB, Beutel R), pp 358-364. deGruyter Press.
  64. Jordal BH, Gebhardt H & Mandelshtam MY. 2013. The red-listed species Thamnurgus rossicus in East Europe is a synonym of the rare Central European species, T. petzi (Curculionidae: Scolytinae). Zootaxa, 3750: 83-88.
  65. Jordal BH. 2013. Deep phylogenetic divergence between Scolytoplatypus and Remansus, a new genus of Scolytoplatypodini from Madagascar (Coleoptera, Curculionidae, Scolytinae). ZooKeys, 352: 9-33.
  66. Jordal BH. 2013. New species and records of Scolytodes (Curclionidae: Scolytinae) from South America. Zootaxa,  3721 (6): 529-551.
  67. Jordal BH & Cognato AI. 2012. Molecular phylogeny of bark and ambrosia beetles reveals multiple origins of fungus farming during periods of global warming. BMC Evolutionary Biology, 12: 133.
  68. Jordal BH. 2012. Phrixosoma concavifrons – a sexually dimorphic Phrixosomatini (Coleoptera: Curculionidae) from the Udzungwa mountains in Tanzania. Zootaxa, 3255: 52-56.
  69. Ruiz CC, Jordal BH & Serrano J. 2012.  Diversification of Calathus (Coleoptera: Carabidae) in the Mediterranean region – glacial refugia and taxon pulses. Journal of Biogeography, 39: 1791-1805.
  70. Andersen HF, Jordal BH, Kambestad M & Kirkendall LR. 2012. Improbable but true: the invasive inbreeding ambrosia beetle Xylosandrus morigerus has generalist genotypes. Ecology and Evolution, 2: 247-257.
  71. Jordal BH, Sequeira AS & Cognato AI. 2011. Age and phylogeny of wood boring weevils and the origin of subsociality. Molecular Phylogenetics and Evolution, 59: 708-724.
  72. Cognato AI, Hulcr J, Dole SA & Jordal BH. 2011. Phylogeny of haplo-diploid, fungus-growing ambrosia beetles (Coleoptera: Curculionidae: Scolytinae: Xyleborini) inferred from molecular and morphological data. Zoologica Scripta, 40: 174-186.
  73. Jordal BH. 2010. Revision of the genus Phloeoditica Schedl - with description of two new genera and two new species in Phloeosinini (Coleoptera: Curculoninidae, Scolytinae). ZooKeys, 56: 141-156.
  74. Dole SA, Jordal BH & Cognato, AI. 2010. Polyphyly of Xylosandrus Reitter inferred from nuclear and mitochondrial genes (Coleoptera: Curculionidae, Scolytinae).  Molecular Phylogenetics and Evolution, 54: 773-782.
  75. Ruiz CC, Jordal BH, Emerson BC & Serrano J.  2009. Molecular phylogeny and Holarctic diversification of the subtribe Calathina (Coleoptera: Carabidae: Sphodrini). Molecular Phylogenetics and Evolution, 55: 358-371.
  76. Jordal BH. 2009.  The Madagascan genus Dolurgocleptes Schedl (Coleoptera: Curculionidae, Scolytinae): description of a new species and transfer to the tribe Polygraphini. Zootaxa, 2014: 41-50.
  77. Jordal BH. 2009.  Two new species of Aphanarthrum (Coleoptera, Scolytinae) associated with Euphorbia in South Africa. Zootaxa, 1983: 54–62.
  78. Ruiz CC, Jordal BH & Serrano J. 2009.  Molecular phylogeny of the tribe Sphodrini (Coleoptera: Carabidae) based on mitochondrial and nuclear markers. Molecular Phylogenetics and Evolution 50:  44-58.
  79. Jordal BH, Gillespie J & Cognato AI. 2008. Secondary structure alignment and direct optimization of 28S nucleotides provide limited phylogenetic resolution in bark and ambrosia beetles (Curculionidae: Scolytinae). Zoologica Scripta, 37: 1-14.
  80. Jordal BH & Knizek M. 2007. Resurrection of Crypturgus subcribrosus Eggers stat. nov. and its close phylogenetic relationship to Nearctic Crypturgus (Curculionidae, Scolytinae). Zootaxa, 1606: 41-50.
  81. Jordal BH. 2007. Reconstructing the phylogeny of Scolytinae and close allies:  major obstacles and prospects for a solution. USDA Forest Service Proceedings RMRS-P-45: 3-8.
  82. Jordal BH, Emerson, BC & Hewitt, GM. 2006. Apparent 'sympatric' speciation in ecologically similar herbivorous beetles facilitated by multiple colonisations of an island. Molecular Ecology, 15: 2935-2947.
  83. Kirkendall LR & Jordal BH. 2006. The bark and ambrosia beetles (Curculionidae, Scolytinae) of Cocos Island, Costa Rica and the role of mating systems in island zoogeography. Biological Journal of the Linnean Society, 89: 729-743 [includes description of three new species].
  84. Jordal BH. 2006. Community structure and reproductive biology in bark beetles (Coleoptera: Scolytinae) associated with Macaronesian Euphorbia spurges. European Journal of Entomology, 103: 71-80.
  85. Jordal BH & Hewitt GM. 2004. The origin and radiation of Macaronesian beetles breeding in Euphorbia:  the relative importance of multiple data partitions and population sampling. Systematic Biology, 53: 711-734. 
  86. Jordal BH, Kirkendall LR & Harkestad K. 2004. Phylogeny of a Macaronesian radiation: host-plant use and possible cryptic speciation in Liparthrum bark beetles. Molecular Phylogenetics and Evolution, 31: 554-571.
  87. Nakagawa M, Itioka T, Momose K. Yumoto T, Komai F, Morimoto K, Jordal BH, Kato M, Kaliang H, Hamid AA, Inoue T & Nakashizuka T. 2003. Resource use of insect seed predators during general flowering and seeding events in a Bornean dipterocarp rain forest. Bulletin of Entomological Research, 93: 455-466.
  88. Jordal BH, Beaver RA, Normark BB & Farrell BD. 2002. Extraordinary sex ratios, and the evolution of male neoteny in sib-mating Ozopemon beetles. Biological Journal of the Linnean Society, 75: 353-360.
  89. Jordal BH. 2002. Elongation Factor 1a  resolves the monophyly of the haplodiploid ambrosia beetles Xyleborini (Coleoptera: Curculionidae). Insect Molecular Biology, 11: 453-465.
  90. Jordal BH, Normark BB, Farrell BD & Kirkendall LR. 2002. Extraordinary haplotype diversity in haplodiploid inbreeders:  phylogenetics and evolution of the bark beetle genus Coccotrypes. MolecularPhylogenetics and Evolution, 23: 171-188.
  91. Jordal BH, Beaver RA & Kirkendall LR. 2001. Breaking taboos in the tropics: inbreeding promotes colonization by wood-boring beetles. Global Ecology and Biogeography 10: 345-358.
  92. Farrell BD, Sequeira A, O'Meara B, Normark BB, Chung J & Jordal BH. 2001. The evolution of agriculture in beetles (Curculionidae: Scolytinae and Platypodinae). Evolution, 55: 2011-2027.
  93. Jordal BH, Normark BB & Farrell BD. 2000. Evolutionary radiation of an inbreeding haplodiploid beetle lineage (Curculionidae, Scolytinae). Biological Journal of the Linnean Society 71: 483-499.
  94. Normark BB, Jordal BH & Farrell BD. 1999. Origin of a haplodiploid beetle lineage. Proceedings of the Royal Society of London serie B 266: 2253-2259.
  95. Jordal BH. 1998. A review of Scolytodes Ferrari (Coleoptera: Scolytidae) associated with Cecropia (Cecropiaceae) in the northern Neotropics. Journal of Natural History 32: 31-84. 
  96. Jordal BH & Kirkendall LR. 1998. Ecological relationships of a guild of tropical beetles breeding in Cecropia leafstalks in Costa Rica. Journal of Tropical Ecology 14: 153-176.
  97. Jordal BH. 1998. New species of Scolytodes (Coleoptera: Scolytidae) from Costa Rica and Panama. Revista de Biologia Tropical 46: 407-420.
  • Bark and ambrosia beetles of Africa and Madagascar - taxonomic revisions
  • Biogeography of Madagascar
  • Revision of Scolytodes bark beetles
  • Norwegian Oribatida mites
  • Museum collection data - GBIF/Artsdatabanken
  • Fylogenetisk analyse av insekter, spesielt biller
  • Evolusjon av gener og hvordan DNA sekvenser fra slike gener kan optimaliseres og benyttes i standardiserte analyser av evolusjonært slektskap.
  • Taksonomi av vedborende biller, spesielt barkbiller.