Fish macro- and microanatmy, in particular the structure and development of the skeletal system.

BIO 280 Fish biology: Anatomy part

BIO 381 Fish histopathology: Normal fish histology

(BIO 270 Vertebrate anatomy: Macroanatomy

BIO 305 Methods in microscopy)

Academic article

• Fjelldal, Per Gunnar; Murugesan, Sankar; Vågseth, Tone et al. (2025). Vertebral deformities in cultured big size Rainbow Trout: Radiological analysis from juvenile to harvest size. (external link)
• Murugesan, Sankar; Fraser, Thomas; Kryvi, Harald et al. (2025). Effect of heterozygosity, ploidy and incubation temperature on post-cranial axial skeletal meristics and deformities in Atlantic salmon (Salmo salar). (external link)
• Philip, Antony Jesu Prabhu; Fraser, Thomas; Nordvik, Kari et al. (2024). Sequence of formation and inheritance of meristic variation in the post-cranial axial skeleton of Atlantic salmon (Salmo salar). (external link)
• Murugesan, Sankar; Kryvi, Harald; Fraser, Thomas et al. (2024). A new method for regionalization of the vertebral column in salmonids based on radiographic hallmarks. (external link)
• Engdal, Vilde Arntzen; Dalum, Alf Seljenes; Kryvi, Harald et al. (2024). State of the heart: Anatomical annotation and assessment of morphological cardiac variation in Atlantic salmon (Salmo salar L.). (external link)
• Takvam, Marius; Denker, Elsa; Gharbi, Naouel et al. (2021). Sulfate homeostasis in Atlantic salmon is associated with differential regulation of salmonid-specific paralogs in gill and kidney. (external link)
• Kryvi, Harald; Nordvik, Kari; Fjelldal, Per Gunnar et al. (2020). Heads and tails: The notochord develops differently in the cranium and caudal fin of Atlantic Salmon (Salmo salar, L.). (external link)
• Kryvi, Harald; Rusten, Iselin; Fjelldal, Per Gunnar et al. (2017). The notochord in Atlantic salmon (Salmo salar L.) undergoes profound morphological and mechanical changes during development. (external link)
• Humborstad, Odd Børre; Ferter, Keno; Kryvi, Harald et al. (2016). Exophthalmia in wild-caught cod (Gadus morhua L.): development of a secondary barotrauma effect in captivity. (external link)
• Fjelldal, Per Gunnar; Solberg, Monica Favnebøe; Hansen, Tom Johnny et al. (2016). Salmonid fish: model organisms to study cardiovascular morphogenesis in conjoined twins?. (external link)
• Eide, Marta; Karlsen, Odd Andre; Kryvi, Harald et al. (2014). Precision-cut liver slices of Atlantic cod (Gadus morhua): An in vitro system for studying the effects of environmental contaminants. (external link)
• Wang, Shou; Furmanek, Tomasz; Kryvi, Harald et al. (2014). Transcriptome sequencing of Atlantic salmon (Salmo salar L.) notochord prior to development of the vertebrae provides clues to regulation of positional fate, chordoblast lineage and mineralisation. (external link)
• Fjelldal, Per Gunnar; Totland, Geir Kåre; Hansen, Tom Johnny et al. (2013). Regional changes in vertebra morphology during ontogeny reflect the life history of Atlantic cod (Gadus morhua L.). (external link)
• Wang, Shou; Kryvi, Harald; Grotmol, Sindre et al. (2013). Mineralization of the vertebral bodies in Atlantic salmon (Salmo salar L.) is initiated segmentally in the form of hydroxyapatite crystal accretions in the notochord sheath. (external link)
• Totland, Geir Kåre; Fjelldal, Per Gunnar; Kryvi, Harald et al. (2011). Sustained swimming increases the mineral content and osteocyte density of salmon vertebral bone. (external link)
• Sagstad, Anita; Grotmol, Sindre; Kryvi, Harald et al. (2011). Identification of vimentin- and elastin-like transcripts specifically expressed in developing notochord of Atlantic salmon (Salmo salar L.). (external link)
• Kamisaka, Yuko; Jordal, Ann-Elise Olderbakk; Edvardsen, Rolf et al. (2010). A case report on the distended gut syndrome (DGS) in cultured larvae of Atlantic cod (Gadus morhua). (external link)
• Marks, Jessica; Biermann, Christiane H.; Eanes, Walter F. et al. (2008). Sperm Polymorphism Within the Sea Urchin Strongylocentrotus droebachiensis: Divergence Between Pacific and Atlantic Oceans. (external link)
• Grotmol, Sindre; Kryvi, Harald; Keynes, Roger et al. (2006). Stepwise enforcement of the notochord and its intersection with the myoseptum: an evolutionary path leading to development of the vertebra?. (external link)
• Grotmol, Sindre; Totland, Geir Kåre; Kryvi, Harald (2005). Deformation of the notochord by pressure from the swim bladder may cause malformation of the spine in reared Atlantic cod larvae: a case study. (external link)
• Nordvik, Kari; Kryvi, Harald; Totland, Geir Kåre et al. (2005). The salmon vertebral body develops through mineralization of two preformed tissues that are encompassed by two layers of bone. (external link)
• Grotmol, Sindre; Nordvik, Kari; Kryvi, Harald et al. (2005). A segmental pattern of alkaline phosphatase activity with the notochord coincides with the initial formation of the vertebrae. (external link)
• Grotmol, Sindre; Nordvik, Kari; Kryvi, Harald et al. (2005). A segmental pattern of alkaline phosphatase activity within the notochord coincides with the initial formation of the vertebral bodies. (external link)
• Fjelldal, PG; Grotmol, Sindre; Kryvi, Harald et al. (2004). Pinealectomy induces malformation of the spine and reduces the mechanical strength of the vertebrae in Atlantic salmon, Salmo salar. (external link)
• Grotmol, Sindre; Kryvi, Harald; Nordvik, Kari et al. (2003). Notochord segmentation may lay down the pathway for the development of the vertebral bodies in the Atlantic salmon. (external link)
• Grotmol, Sindre; Kryvi, Harald; Nordvik, Kari et al. (2003). Notochord segmentation may lay the pathway for the development of the vertebral bodies of the Atlantic salmon Salmo salar. (external link)
• Grotmol, Sindre; Totland, Geir Kåre; Kryvi, Harald et al. (2002). Spatial distribution of muscle fibre types within skeletal muscle fascicules from standardbred horses. (external link)
• Grotmol, Sindre; Totland, Geir Kåre; Kryvi, Harald et al. (2002). Spatial distribution of fiber types within skeletal muscle fascicles from Standardbred horses. (external link)
• Ringø, Einar; Mayhew, Terry; Kryvi, Harald et al. (2001). Damaging effect of dietary inulin to intestinal enterocytes in Arctic charr (Salvelinus alpinus L.). (external link)
• Ringø, Einar; Mayhew, Terry; Kryvi, Harald et al. (2001). Damaging effect of dietary inulin to intestinal enterocytes in Arctic charr (Salvelinus alpinus L.). (external link)
• Totland, Geir Kåre; Madsen, Lise; Klementsen, Beate et al. (2000). Proliferation of mitochondria and gene expression of carnitine palmitoyltransferase and fatty acyl-CoA oxidase in rat skeletal muscle, heart and liver by hypolipidemic fatty acids. (external link)
• Taranger, G.O.; Haux, C.; Stefansson, S.O. et al. (1999). Mechanisms underlying photoperiodic effects on age at sexual maturity in Atlantic salmon, Salmo salar L. (external link)
• Taranger, Geir Lasse; Haux, C.; Hansen, Tom Johnny et al. (1999). Mechanisms underlying photoperiodic effects on age at sexual maturity in Atlantic salmon, Salmo salar. (external link)
• Galloway, T. F.; Kjørsvik, E.; Kryvi, Harald (1998). Effect of temperature and viability and axial muscle development in embryos and yolk sac larvae of the northeast arctic cod (Gadus morhua). (external link)
• Frøyland, Livar; Helland, Knut; Kryvi, Harald et al. (1996). Increased fatty acid oxidation and polydispersity of rat liver mitochondria and peroxisomes induced by the hypolipidemic agent tetradecylthioacetic acid. (external link)
• Kjesbu, Olav Sigurd; Kryvi, Harald; Norberg, Birgitta (1996). Oocyte size and structure in relation to blood plasma steroid hormones in individually monitored, spawning Atlantic cod. (external link)
• Frøyland, Livar; Helland, Knut; Totland, Geir Kåre et al. (1996). A hypolipidemic peroxisome proliferating fatty acid induces polydispersity of rat liver mitochondria. (external link)
• Nissling, A.; Kryvi, Harald; Valin, L. (1994). Variation in egg bouancy of baltic cod (Gadus morhua) and implications for egg survival at prevailing conditions in the Baltic sea. (external link)
• Hamre, K.; Hjeltnes, B.; Kryvi, Harald et al. (1994). Decreased concentration of haemoglobin, accumulation of lipid oxidation products and unchanged skeletal muscle in Atlantic salmon (Salmo salar) fed low dietary vitamin E. (external link)
• Hamre, K.; Hjeltnes, B.; Kryvi, Harald et al. (1994). Decreased concentration of hemaglobin, accumulation of lipid oxidation products and unchanged skeletal muscle in atlantic salmon (Salmo salar) fed on low diatary vitamin E. (external link)
• Hamre, K.; Hjeltnes, B.; Kryvi, Harald et al. (1994). Decreased concentration of hemoglobin, accumulation of lipid oxidation products and unchanged skeletal muscle in atlantic salmon (Salmo salar) fed on low diatary viatamin E. (external link)
• Hamre, K.; Hjeltnes, B.; Kryvi, Harald et al. (1994). Decreased concentration of Hemoglobin, accumualtion of lipid oxidation products and unchanged skeletal muscle in Atlantic salmon (Salmo salar) fed low dietary vitamin E. (external link)
• Kjesbu, Olav Sigurd; Klungsøyr, Jarle; Witthames, Peter R. et al. (1991). Fecundity, atresia, and egg size of captive Atlantic cod (Gadus morhua) in relation to proximate body composition. (external link)
• Kryvi, Harald; Aarsland, A.; Berge, Rolf Kristian (1990). Morphologic effects of sulfursubstituted fatty acids on rat hepatocytes with special references to proliferation of peroxisomes and mitochondria. (external link)
• Slinde, Erik; Baardseth, Pernille; Kryvi, Harald (1983). Effect of ascorbat on a hypotenically isolated mitochondrial fraction during ageing of Swede (Brassica napus var. napobrassica L). (external link)
• Slinde, Erik; Andersen, Knut-Jan; Mørkrid, Lars et al. (1976). Sedimentation Coefficient, Buoyant Density, Morphology, and Optimal Recorvery of Mitochondria from the Ileal Mucosa of Guinea Pigs. (external link)

Lecture

• Takvam, Marius; Sundell, Kristina; Sundh, Henrik et al. (2023). Modifikasjoner av Na+, K+ - ATPase enzymaktivitets protokollen og bruksområder i moderne lakseoppdrett. (external link)
• Kryvi, Harald (1996). Anathomy and phylogeny of the carps. (external link)

Academic literature review

• Takvam, Marius; Wood, Chris M.; Kryvi, Harald et al. (2023). Role of the kidneys in acid-base regulation and ammonia excretion in freshwater and seawater fish: implications for nephrocalcinosis. (external link)
• Takvam, Marius; Sundell, Kristina; Sundh, Henrik et al. (2023). New wine in old bottles: Modification of the Na+/K+-ATPase enzyme activity assay and its application in salmonid aquaculture. (external link)
• Takvam, Marius; Wood, Chris M.; Kryvi, Harald et al. (2021). Ion transporters and osmoregulation in the kidney of teleost fishes as a function of salinity. (external link)

Academic lecture

• Wang, Shou; Furmanek, Tomasz; Kryvi, Harald et al. (2013). DEEP SEQUENCING REVEALS MINERALIZING NOTOCHORD TRANSCRIPTOME, DURING AXIAL SKELETON ENFORCEMENT FROM NOTOCHORD TO VERTEBRA IN ATLANTIC SALMON (SALMO SALAR L.). (external link)
• Wang, Shou; Furmanek, Tomasz; Kryvi, Harald et al. (2012). The Atlantic salmon (Salmo Salar L.) notochord displays a dynamic transcriptome prior to and during vertebral segmentation. (external link)
• Lock, Erik-Jan; Krossøy, Christel; Waagbø, Rune et al. (2008). Roles of fat soluble vitamins in bone development and mineral metabolism. (external link)
• Ørnsrud, Robin; Krossøy, Christel; Waagbø, Rune et al. (2006). Fettløselige vitaminers rolle i beinutvikling og mineralmetabolisme. (external link)
• Ørnsrud, Robin; Grotmol, Sindre; Kryvi, Harald et al. (2004). Roles of fat soluble vitamins in bone development and mineral metabolism. (external link)
• Flood, P. R.; Hjeltnes, B.; Totland, Geir Kåre et al. (1994). Tidlig smittespredning fra ILA-infisert fisk. (external link)
• Søvik, Oddmund; Monsen, A. L. B.; Berge, Rolf Kristian et al. (1994). Zellweger-like syndrome: Prolonged survival in male twins during treatment with n-3 fatty acids. (external link)
• Hjeltnes, B.; Flood, P. R.; Totland, Geir Kåre et al. (1994). Transmission of infectious salmon anemia (ISA) throug naturally secreted material. (external link)
• Totland, Geir Kåre; Hjeltnes, B.; Flood, P. R. et al. (1994). Naturlige smitteveier for ILA. (external link)

Popular scientific article

• Kamisaka, Yuko; Jordal, Ann-Elise Olderbakk; Edvardsen, Rolf et al. (2010). Tarmlidelser hos torskelarver– hvor ligger forklaringene?. (external link)
• Grotmol, Sindre; Cieplinska, Teresa; Kryvi, Harald et al. (2003). "Nakkeknekk" hos torskeyngel. (external link)
• Kryvi, Harald (1996). Kjevebevegelsen hos fisk. (external link)

Masters thesis

• Drønnesund, Egil-André; Kryvi, Harald; Saraste, Jaakko (2006). Distribution of markers of early secretory compartments in pancreatic exocrine cells. (external link)
• Totland, Geir Kåre; Kryvi, Harald; Grotmol, Sindre et al. (2006). Morfologiske effekter av vaksineindusert bukhinnebetennelse på bukspyttkjertelen hos Atlanterhavslaks (Salmo salar L.). (external link)
• Aarskog, Nina Karin; Doskeland, Stein Ove; Kryvi, Harald (1992). Effects of cAMP elevating agents and toxins on differentiation and viability of cultured neuroblastoma cells.. (external link)

Poster

• Waagbø, Rune; Breck, Olav; Kryvi, Harald et al. (2006). Workshop innen beindeformiteter hos laks og torsk i intensivt oppdrett. (external link)
• Skjærven, Kaja H; Finn, Roderick Nigel; Kryvi, Harald et al. (2002). Yolk resorption in developing plaice (Pleuronectes platessa). (external link)

Academic chapter/article/Conference paper

• Skjærven, Kaja H; Finn, Roderick Nigel; Kryvi, Harald et al. (2003). Yolk resorption in developing plaice (Pleuronectes platessa). (external link)
• Kryvi, Harald; Lysebo, H. (1995). Higher education in biology in Norway. (external link)
• Kristensen, I.; Kryvi, Harald; Mortensen, S. (1994). Sjøpattedyr. (external link)
• Kryvi, Harald; Mortensen, S. (1994). Fiskenes anatomi og fysiologi. (external link)
• Berge, Rolf Kristian; Kryvi, Harald; Aarsæther, Niels et al. (1993). A review in peroxisomes, biology and importance in toxicology and medicine. (external link)
• Lossius, Ivar; Kryvi, Harald; Lorens, James et al. (1993). Cloning and characterization of growth regulating genes from Atlantic salmon: A strategy for increasing growth rate by microinjection of a modified growth hormone gene into fertilized salmon eggs. (external link)
• Berge, Rolf Kristian; Kryvi, Harald; Aarsæther, Niels et al. (1993). Proliferation of peroxisomes and mitochondria and modulation of lipid metabolizing enzymes by hypolipidaemic non-_b-oxidizable fatty acids (3-thia fatty acids). (external link)
• Flatmark, Torgeir; Kvannes, Jon; Eikhom, T. S. et al. (1992). Differential responses at the peroxisomal level in hepatocytes and myocardium on feeding rats diets containing partially hydrogenated fish oil. (external link)

Compendium

• Totland, G.K.; Kryvi, Harald (1998). VERTEBRATENES ANATOMI. Kompendium for grunnemne i biologi ved UiB. (external link)

Non-fiction book

• Kryvi, Harald; Totland, Geir Kåre (1995). Fiskeanatomi. (external link)

Doctoral dissertation

• Kryvi, Harald (1978). Studies on the histochemistry, ultrastructure and growth of axial muscle fibres in the sharks Etmopterus spinax and Galeus melastomus. (external link)

See a complete overview of publications in Cristin.

See the list for the research group.

See the list for the research group.