Research
Biostatistics
I am interested in applications of statistics and probability to biology, especially to marine ecology. I have developed methods for line transect surveys and CKMR (close-kin mark recapture) methods in particular. Key reference: Bravington, MV, Skaug, HJ, Anderson, EC (2016). Close-kin Mark-Recapture. Statistical Science.
Computational statistics
My opinion for a long time has been that Statisticians should have Automatic Differentiation (AD) in their toolbox. My conjecture has been that a combination of AD and the Laplace approximation is a powerful platform for fitting models with continuous latent variables. Software projects in which this idea is implemented are:
- ADMB: http://admb-project.org/
Artificial intelligence (AI)
Witnessing the recent breakthroughs in the field, my research interests have turned towards AI. Techniques such as variational autoencoders and diffusion models are probabilistic in nature, and Backpropagation in Deep Learning is the same computational technique as AD, which I have been working with for 20 years.
Editorial services
co-Editor in chief of Scandinavian Journal of Statistics (2018–2021)
Teaching
Teaching at UiB (last 5 years)
Short courses / workshops (in the past)
- CKMR (Close-kin mark recapture) course (co-thought) Dalhousie University, Halifax, July 2019.
- TMB (Template Model Builder) course (co-thought) at IMR Bergen, June 2019.
- Modelling in TMB (Template Model Builder). Course (co-thought) at IMR Bergen, June 2018.
- An introduction to fish stock assessment in TMB (Template Model Builder). Course given Oct. 2017 in La Jolla
Publications
From CRIStin (Current research information system in Norway):
2024
- Førland, Brage; Skaug, Hans Julius; Takahashi, Megumi et al. (2024). Close-Kin Mark-Recapture Without Age Information. (external link)
- Takahashi, Megumi; Førland, Brage; Pastene, Luis A. et al. (2024). Geographical distribution of close kin in southern right whales on feeding grounds. (external link)
- Helgøy, Ingvild Margrethe; Skaug, Hans Julius; Li, Yushu (2024). Sparse Bayesian learning using TMB (Template Model Builder). (external link)
2022
- Delaval, Aurelien Nicolas; Bendall, Victoria; Hetherington, Stuart J. et al. (2022). Evaluating the suitability of close-kin mark-recapture as a demographic modelling tool for a critically endangered elasmobranch population. (external link)
- Andersson, Jonas; Nilsen, Øivind A; Skaug, Hans Julius (2022). Mixed Frequency Data in a (S,s) Pricing Model. (external link)
- Azzolini, Francesca; Berentsen, Geir Drage; Skaug, Hans Julius et al. (2022). The heritability of BMI varies across the range of BMI—a heritability curve analysis in a twin cohort. (external link)
2021
- Mannseth, Janne; Berentsen, Geir Drage; Skaug, Hans Julius et al. (2021). Variation in use of Caesarean section in Norway: An application of spatio-temporal Gaussian random fields. (external link)
- Wacker, Sebastian; Skaug, Hans Julius; Forseth, Torbjørn et al. (2021). Considering sampling bias in close-kin mark–recapture abundance estimates of Atlantic salmon. (external link)
- Nilsen, Geir Kjetil; Munthe-Kaas, Antonella Zanna; Skaug, Hans Julius et al. (2021). Epistemic uncertainty quantification in deep learning classification by the Delta method. (external link)
- Smout, Sophie; Murray, Kimberly; Aarts, Geert et al. (2021). Report of the NAMMCO-ICES Workshop on Seal Modelling (WKSEALS 2020). (external link)
- Solvang, Hiroko Kato; Skaug, Hans Julius; Øien, Nils Inge (2021). Measurement error model for the Norwegian common minke whale (Balaenoptera acutorostrata acutorostrata) surveys 2008-2013. (external link)
- Helgøy, Ingvild Margrethe; Skaug, Hans Julius (2021). The Sibling Distribution for Multivariate Life Time Data. (external link)
- Solvang, Hiroko Kato; Skaug, Hans Julius; Øien, Nils Inge (2021). Consideration of measurement errors for the Norwegian common minke whale (Balaenoptera acutorostrata acutorostrata) surveys. (external link)
2020
- Berentsen, Geir Drage; Azzolini, Francesca; Skaug, Hans J. et al. (2020). Heritability curves: A local measure of heritability in family models. (external link)
- Frøysa, Håvard G; Skaug, Hans J.; Alendal, Guttorm (2020). Experimental design for parameter estimation in steady-state linear models of metabolic networks. (external link)
- Frøysa, Håvard G; Alendal, Guttorm; Skaug, Hans J. et al. (2020). Estimation and identifiability of kinetic parameters in dynamical models of biochemical reaction networks. (external link)
- Fallahi, Shirin; Skaug, Hans J.; Alendal, Guttorm (2020). A comparison of Monte Carlo sampling methods for metabolic network models. (external link)
2019
- Nikolioudakis, Nikolaos; Skaug, Hans J.; Olafsdottir, Anna H. et al. (2019). Drivers of the summer-distribution of Northeast Atlantic mackerel (Scomber scombrus) in the Nordic Seas from 2011 to 2017; a Bayesian hierarchical modelling approach.. (external link)
- Dale, Karina; Müller, Mette Bjørge; Tairova, Zhanna et al. (2019). Contaminant accumulation and biological responses in Atlantic cod (Gadus morhua) caged at a capped waste disposal site in Kollevåg, Western Norway. (external link)
- Gjessing, Håkon K.; Skaug, Hans J. (2019). Editorial. (external link)
- Mannseth, Janne; Skaug, Hans J. (2019). Bayesian and frequentist computation with application to data from the Medical Birth Registry of Norway. (external link)
- Skaug, Hans J. (2019). Integration by differentiation; almost a free lunch. (external link)
- Ruzzante, Daniel E.; McCracken, Gregory R; Førland, Brage et al. (2019). Validation of close‐kin mark–recapture (CKMR) methods for estimating population abundance. (external link)
- Skaug, Hans J. (2019). Close-Kin Mark-Recapture (CKMR). (external link)
- Skaug, Hans J.; Bravington, Mark V. (2019). Close kin mark recapture in fisheries . (external link)
- Skaug, Hans J. (2019). Close-Kin Mark-Recapture (CKMR). (external link)
- Shah, Abhishek; Bertino, Laurent; Skaug, Hans J. (2019). Stochastic data assimilation of observations with a detection limit. (external link)
- Nilsen, Geir Kjetil; Brun, Morten; Zanna, Antonella et al. (2019). Efficient Computation of Hessian Matrices in TensorFlow . (external link)
- Goksøyr, Anders; Alendal, Guttorm; Aranguren-Abadía, Libe et al. (2019). dCOD 1.0: DECODING THE SYSTEMS TOXICOLOGY OF ATLANTIC COD (GADUS MORHUA) – HIGHLIGHTS SO FAR. (external link)
- Goksøyr, Anders; Alendal, Guttorm; Aranguren-Abadía, Libe et al. (2019). dCOD 1.0: DECODING THE SYSTEMS TOXICOLOGY OF ATLANTIC COD (GADUS MORHUA) – HIGHLIGHTS SO FAR. (external link)
- Fallahi, Shirin; Skaug, Hans J.; Alendal, Guttorm (2019). Bayesian flux balance analysis of metabolic pathways. (external link)
2018
- Frøysa, Håvard G; Skaug, Hans J.; Alendal, Guttorm (2018). Experimental design for estimation of first order kinetic parameters in steady-state metabolic networks. (external link)
- Fallahi, Shirin; Skaug, Hans J.; Alendal, Guttorm (2018). Bayesian flux balance analysis; concepts and computation. (external link)
- Gundersen, Kristian; Oleynik, Anna; Alendal, Guttorm et al. (2018). Model Reduction for Tracer Transport and Applications. (external link)
- Oleynik, Anna; Gundersen, Kristian; Alendal, Guttorm et al. (2018). Mathematical methods for detection and localization of CO2 leaks. (external link)
- Dale, Karina; Karlsen, Odd Andre; Yadetie, Fekadu et al. (2018). Ecotoxicological responses in Atlantic cod (Gadus morhua) after caging at a capped waste disposal site in Kollevåg, Western Norway . (external link)
- Goksøyr, Anders; Alendal, Guttorm; Aranguren Abadía, Libe et al. (2018). dCOD 1.0: DECODING THE SYSTEMS TOXICOLOGY OF ATLANTIC COD (GADUS MORHUA). (external link)
- Dale, Karina; Karlsen, Odd Andre; Yadetie, Fekadu et al. (2018). Ecotoxicological responses in Atlantic cod (Gadus morhua) after caging at a capped waste disposal site in Kollevåg, Western Norway . (external link)
- Gundersen, Kristian; Oleynik, Anna; Alendal, Guttorm et al. (2018). Combining Models and Machine Learning Techniques to Design Leak Detection Monitoring . (external link)
- Gundersen, Kristian; Oleynik, Anna; Skaug, Hans J. et al. (2018). Machine Learning in CO2 leak detection. (external link)
2017
- Yadetie, Fekadu; Eide, Marta; Söderström, Sofie et al. (2017). PRECISION-CUT LIVER SLICES (PCLS) FOR TOXICOGENOMIC STUDIES IN ATLANTIC COD (Gadus morhua): THE dCOD 1.0 PROJECT. (external link)
- Gundersen, Kristian; Blaser, Nello; Alendal, Guttorm et al. (2017). Assessment of Topological Data Analysis and Machine Learning Technologies as Tools for Seep Detection. (external link)
- Alendal, Guttorm; Blackford, Jeremy; Chen, Baixin et al. (2017). Bayes’ theorem as the fundament to design monitoring programs.. (external link)
- Fallahi, Shirin; Kleppe, Rune; Goksøyr, Anders et al. (2017). Mathematical modeling of regulatory interactions in the fatty acid synthesis pathway. (external link)
- Frøysa, Håvard G; Skaug, Hans J.; Alendal, Guttorm (2017). Structural identifiability of kinetic parameters. (external link)
- Fallahi, Shirin; Kleppe, Rune; Goksøyr, Anders et al. (2017). Mathematical modeling of regulatory interactions in the fatty acid synthesis pathway. (external link)
- Fallahi, Shirin; Kleppe, Rune; Goksøyr, Anders et al. (2017). Mathematical modeling of regulatory interactions in the fatty acid synthesis pathway. (external link)
- Kristensen, Kasper; Berentsen, Geir Drage; Brooks, Mollie et al. (2017). TMB: a flexible framework for developing mixed model software in R. (external link)
- Malde, Ketil; Seliussen, Bjørghild Breistein; Sanchez, Maria Quintela et al. (2017). Whole genome resequencing reveals diagnostic markers for investigating global migration and hybridization between minke whale species. (external link)
- Brooks, Mollie E.; Kristensen, Kasper; van Benthem, Koen J. et al. (2017). glmmTMB balances speed and flexibility among packages for zero-inflated generalized linear mixed modeling. (external link)
- Skaug, Hans J. (2017). The parent-offspring probability when sampling age-structured populations. (external link)
- Goksøyr, Anders; Celander, Malin; Alendal, Guttorm et al. (2017). dCod 1.0: decoding the systems toxicology of Atlantic cod (Gadus morhua). (external link)
- Gundersen, Kristian; Alendal, Guttorm; Skaug, Hans J. et al. (2017). Assessment of machine learning methods as a tool in detecting leakages. (external link)
- Frøysa, Håvard G; Skaug, Hans J.; Alendal, Guttorm (2017). Structural identifiability of kinetic parameters. (external link)
- Goksøyr, Siri Øfsthus; Goksøyr, Anders; Karlsen, Odd Andre et al. (2017). dCod 1.0 decoding the systems toxicology of Atlantic cod. (external link)
- Mannseth, Janne; Kleppe, Tore Selland; Skaug, Hans J. (2017). On the application of improved symplectic integrators in Hamiltonian Monte Carlo. (external link)
2016
- Alendal, Guttorm; Blackford, Jeremy; Chen, Baixin et al. (2016). Bayes’ theorem as the fundament to design monitoring programs.. (external link)
- Kristensen, Kasper; Nielsen, Anders; Berg, Casper W. et al. (2016). TMB: Automatic differentiation and laplace approximation. (external link)
- Vincenzi, Simone; Crivelli, Alain J.; Munch, Stephan et al. (2016). Trade-offs between accuracy and interpretability in von Bertalanffy random-effects models of growth. (external link)
- Bravington, Mark V.; Skaug, Hans J.; Anderson, Eric C. (2016). Close-kin mark-recapture. (external link)
- Lunde, Berent Ånund Strømnes; Skaug, Hans J. (2016). Likelihood Estimation of Jump-Diffusions: Extensions from Diffusions to Jump-Diffusions, Implementation with Automatic Differentiation, and Applications. (external link)
2015
- Zhang, Yanhui; Oliver, Dean; Chen, Yan et al. (2015). Data assimilation by use of the iterative ensemble smoother for 2D facies models. (external link)
- Andersson, Jonas; Nilsen, Øivind Anti; Skaug, Hans J. (2015). Panel data model with mixed frequency – an application to producers' pricing behaviour. (external link)
- Øigård, Tor Arne; Skaug, Hans J. (2015). Fitting state-space models to seal populations with scarce data. (external link)
- Thorson, James T.; Shelton, Andrew O.; Ward, Eric J. et al. (2015). Geostatistical delta-generalized linear mixed models improve precision for estimated abundance indices for West Coast groundfishes. (external link)
- Thorson, James T.; Scheuerell, Mark D.; Shelton, Andrew O. et al. (2015). Spatial factor analysis: A new tool for estimating joint species distributions and correlations in species range. (external link)
- Kleppe, Tore Selland; Skaug, Hans J. (2015). Bandwidth selection in pre-smoothed particle filters. (external link)
- Thorson, James T.; Skaug, Hans J.; Kristensen, Kasper et al. (2015). The importance of spatial models for estimating the strength of density dependence. (external link)
- Oedekoven, Cornelia S; Laake, Jeffrey L; Skaug, Hans J. (2015). Distance sampling with a random scale detection function. (external link)
- Haug, Tore; Øien, Nils Inge; Skaug, Hans J. et al. (2015). DNA-register for vågehval: en skattekiste for studier av hvalbiologi.. (external link)
- Solvang, Hiroko Kato; Skaug, Hans J.; Øien, Nils Inge (2015). Measurement error model for the Norwegian minke whale surveys 2008-2013. (external link)
- Solvang, Hiroko Kato; Skaug, Hans J.; Øien, Nils Inge (2015). Abundance estimates of common minke whales in the Northeast Atlantic base don survey data collected over the period 2008-2013. (external link)
2014
- Anderson, Eric C.; Skaug, Hans J.; Barshis, David J. (2014). Next-generation sequencing for molecular ecology: a caveat regarding pooled samples. (external link)
- Lorentzen, Rolf; Stordal, Andreas S.; Nævdal, Geir et al. (2014). Estimation of production rates with transient well-flow modeling and the auxiliary particle filter. (external link)
- Skaug, Hans J.; Yu, Jun (2014). A flexible and automated likelihood based framework for inference in stochastic volatility models. (external link)
- Vincenzi, Simone; Mangel, Marc; Crivelli, Alain J. et al. (2014). Determining individual variation in growth and its implication for life-history and population processes using the Empirical Bayes Method. (external link)
- Kleppe, Tore Selland; Yu, Jun; Skaug, Hans J. (2014). Maximum likelihood estimation of partially observed diffusion models. (external link)
- Sanchez, Maria Quintela; Skaug, Hans J.; Øien, Nils Inge et al. (2014). Investigating population genetic structure in a highly mobile marine organism: The minke whale balaenoptera acutorostrata acutorostrata in the north east atlantic. (external link)
- Solvang, Hiroko Kato; Skaug, Hans J.; Øien, Nils Inge (2014). Preliminary abundance estimates of common minke whales in the Northeast Atlantic based on survey data collected over the period 2008-2013. (external link)
- Kleppe, Tore Selland; Skaug, Hans J. (2014). Bandwidth Selection In Pre-Smoothed Particle Filters. (external link)
- Haug, Tore; Øien, Nils Inge; Skaug, Hans J. et al. (2014). Unødvendig med fem fangstområder for vågehval.. (external link)
2013
- Langrock, Roland; Borchers, David L.; Skaug, Hans J. (2013). Markov-modulated nonhomogeneous poisson processes for modeling detections in surveys of marine mammal abundance. (external link)
- Glover, Kevin; Kanda, Naohisa; Haug, tore et al. (2013). Hybrids between common and Antarctic minke whales are fertile and can back-cross. (external link)
- Haaland, Øystein Ariansen; Skaug, Hans J. (2013). Estimating genotyping error rates from parent-offspring dyads. (external link)
- Bolker, Benjamin M.; Gardner, Beth; Maunder, Mark et al. (2013). Strategies for fitting nonlinear ecological models in R, AD Model Builder, and BUGS. (external link)
2012
- Fournier, David A.; Skaug, Hans J.; Ancheta, Johnoel et al. (2012). AD Model Builder: using automatic differentiation for statistical inference of highly parameterized complex nonlinear models. (external link)
- Okamura, Hiroshi; Minamikawa, Shingo; Skaug, Hans J. et al. (2012). Abundance estimation of long-diving animals using line transect methods. (external link)
- Stordal, Andreas S.; Valestrand, Randi; Karlsen, Hans A et al. (2012). Comparing the adaptive Gaussian mixture filter with the ensemble Kalman filter on synthetic reservoir models. (external link)
- Glover, Kevin Alan; Haug, tore; Øien, Nils Inge et al. (2012). The Norwegian minke whale DNA register: a data base monitoring commercial harvest and trade of whale products. (external link)
- Stordal, Andreas S.; Karlsen, Hans A; Nævdal, Geir et al. (2012). Filtering with state space localized Kalman gain. (external link)
- Torrissen, Ole; Glover, Kevin; Haug, tore et al. (2012). Good ethics or political and cultural censoring in science?. (external link)
- Fossum, Kristian; Mannseth, Trond; Oliver, Dean et al. (2012). Numerical comparison of Ensemble Kalman Filter and Randomized maximum Likelihood. (external link)
- Zhang, Yanhui; Oliver, Dean; Chen, Yan et al. (2012). Data Assimilation Using the Iterative Ensemble Smoother for 2-D Categorical Models. (external link)
- Kleppe, Tore Selland; Skaug, Hans J. (2012). Fitting general stochastic volatility models using Laplace accelerated sequential importance sampling. (external link)
2011
- Haaland, Øystein; Glover, Kevin A.; Seliussen, Bjørghild Breistein et al. (2011). Genotyping errors in a calibrated DNA register: implications for identification of individuals. (external link)
- Thulin, Kristian; Nævdal, Geir; Skaug, Hans Julius et al. (2011). Quantifying Monte Carlo Uncertainty in the Ensemble Kalman Filter. (external link)
- Stordal, Andreas S.; Karlsen, Hans A; Nævdal, Geir et al. (2011). Bridging the ensemble Kalman filter and particle filters: the adaptive Gaussian mixture filter. (external link)
2010
- Glover, Kevin; Kanda, Naohisa; Haug, Tore et al. (2010). Migration of Antarctic Minke Whales to the Arctic. (external link)
- Skaug, Hans Julius; Øien, Nils Inge; Haug, Tore (2010). Hva kan et register over vågehvalens DNA fortelle om adferd og biologi?. (external link)
- Kleppe, Tore Selland; Skaug, Hans J.; hiroshi, okamura (2010). Asymptotic bias of the hazard probability model under model mis-specification. (external link)
- Kleppe, Tore Selland; Green, William; Yu, Jun et al. (2010). Maximum Simulated Likelihood Methods and Applications. (external link)
- Kleppe, Tore Selland; Yu, Jun; Skaug, Hans Julius (2010). Simulated maximum likelihood estimation of continuous time stochastic volatility models. (external link)
- Liow, Lee Hsiang; Skaug, Hans Julius; Ergon, Torbjørn H. et al. (2010). Global occurrence trajectories of microfossils: environmental volatility and the rise and fall of individual species. (external link)
- Økland, Jan-Magnus; Haaland, Øystein; Skaug, Hans Julius (2010). A method for defining management units based on genetically determined close relatives. (external link)
- Skaug, Hans Julius; Berube, Martine; Palsbøll, Per (2010). Detecting dyads of related individuals in large collections of DNA-profiles by controlling the false discovery rate. (external link)
- Le Rouzic, Arnaud; Skaug, Hans Julius; Hansen, Thomas F. (2010). Estimating genetic architectures from artificial-selection responses: A random-effect framework. (external link)
2008
- Kleppe, Tore Selland; Skaug, Hans Julius (2008). Building and Fitting Non-Gaussian Latent Variable Models via the Moment-Generating Function. (external link)
- Paulsen, Jostein; Lunde, Astrid; Skaug, Hans Julius (2008). Fitting mixed-effects models when data are left truncated. (external link)
- Skaug, Hans Julius; Berube, Martine; Rew, Mary Beth et al. (2008). Genetic analyses reveal promiscuous mating in female minke whales, Balaenoptera acutorostrata. (external link)
- Ergon, Torbjørn; Skaug, Hans Julius (2008). Estimating distributions of correlated individual life-history traits from mark-recapture data. (external link)
2006
- Kitakado, T; Kitada, S; Kishino, H et al. (2006). An integrated-likelihood method for estimating genetic differentiation between populations. (external link)
- Skaug, Hans Julius; Fournier, David (2006). Automatic Approximation of the Marginal Likelihood in non-Gaussian Hierarchical Models. (external link)
- Skaug, Hans Julius (2006). Markov modulated Poisson processes for clustered line transect data. (external link)
- Palsbøll, Per; Berube, Martine; Skaug, Hans Julius et al. (2006). DNA registers of legally obtained wildlife and derived products as means to identify illegal takes. (external link)
- Frimannslund, Lennart; Steihaug, Trond; Skaug, Hans Julius (2006). On Curvature and Separability in Unconstrained Optimisation. (external link)
2005
- Skaug, Hans Julius; Øien, Nils (2005). Genetic tagging of male North Atlantic minke whales through comparison of maternal and foetal DNA-profiles. (external link)
- Skaug, Hans Julius; Øien, Nils (2005). Genetic tagging of males in North Atlantic minke whales through comparison of mother and fetus DNA-profiles. (external link)
- Trenkel, Verena; Skaug, Hans Julius (2005). Disentangling the effects of capture efficiency and population abundance on catch data using random effects models. (external link)
2001
1999
1997
- Skaug, Hans J.; Gjøsæter, Harald; Haug, tore et al. (1997). Do Minke Whales (Balaenoptera acutorostrara) Exhibit Particular Prey Preferences?. (external link)
- Schweder, Tore; Skaug, Hans J.; Dimakos, Xeni Kristine et al. (1997). Abundance of northeastern Atlantic minke whales, estimates for 1989 and 1995. (external link)
1996
- Skaug, Hans Julius; Dimakos, Xeni Kristine; Schweder, Tore (1996). NAILS-96 Specification document. (external link)
- Haug, Tore; Lindstrøm, Ulf; Nilssen, Kjell T. et al. (1996). Diet and food avaliability for Northeastern Atlantic minke whales, Balaenoptera Acutorostrata. (external link)
- Belsnes, Dag; Bergan, Marius; Dimakos, Xeni Kristine et al. (1996). NAILS-96, System Design and Program Documentation. (external link)
- Skaug, Hans Julius (1996). Perpendicular distance line transect methods based on integrated spatial hazard probability models. (external link)
- Schweder, Tore; Skaug, Hans Julius; Dimakos, Xeni Kristine et al. (1996). Abundance of northeastern Atlantic minke whales, estimates for 1989 and 1995. (external link)
1994
1993
- Skaug, Hans Julius (1993). The limit distribution of the Hoeffding statistic for test of serial independence. Report No.22 Jan. 1993. (external link)
- Skaug, Hans Julius; Tjøstheim, Dag Bjarne (1993). A nonparametric test of serial independence based on the empirical distribution function. (external link)