Academic article

• Erwan Lagadec; Enrique Pino Martinez; Heidrun Nylund et al. (2026). Temporal dynamics of Tenacibaculum maritimum and Tenacibaculum dicentrarchi in a land-based Atlantic salmon (Salmo salar) facility in Norway. (external link)
• Pablo Balseiro Vigo; Erlend Andreas Nordvoll Holtan; Sara Calabrese et al. (2025). Performance of Atlantic salmon post-smolts reared in RAS with nanofiltered sulfate-reduced brackish water. (external link)
• Floriana Lai; Sergei Budaev; Ida Kristine Hundven et al. (2025). Influence of water temperature on feed intake, appetite control, and energy allocation in Atlantic salmon (Salmo salar) post-smolt. (external link)
• S Fivelstad; AB Olsen; Sigurd Stefansson et al. (2004). Lack of long-term sublethal effects of reduced freshwater pH alone on Atlantic salmon (Salmo salar) smolts subsequently transferred to seawater. (external link)
• Lene Sveen; Gerrit Timmerhaus; Aleksei Krasnov et al. (2018). High fish density delays wound healing in Atlantic salmon (Salmo salar). (external link)
• Trine Ytrestøyl; Harald Rune Takle; Jelena Kolarevic et al. (2020). Performance and welfare of Atlantic salmon, Salmo salar L. post-smolts in recirculating aquaculture systems: Importance of salinity and water velocity. (external link)
• Enrique Pino Martinez; Pablo Balseiro; Mitchell S. Fleming et al. (2023). Interaction of temperature and photoperiod on male postsmolt maturation of Atlantic salmon (Salmo salar L.). (external link)
• Gunnar Steinn Gunnarsson; Egil Karlsbakk; Steffen H. Blindheim et al. (2017). Temporal changes in infections with some pathogens associated with gill disease in farmed Atlantic salmon (Salmo salar L). (external link)
• Camilla Diesen Hosfeld; Sigurd O Handeland; Sveinung Fivelstad et al. (2010). Physiological effects of normbaric environmental hyperoxia on Atlantic salmon (Salmo solar L.) presmolts. (external link)
• Kristin Kvamme; Sveinung Fivelstad; Sigurd O Handeland et al. (2018). Water flow and diurnal variation in metabolite production rates of Atlantic salmon (Salmo salar L.) post-smolt. (external link)
• Rouzbeh Keihani; Ana S. Gomes; Pablo Balseiro Vigo et al. (2024). Evaluation of stress in farmed Atlantic salmon (Salmo salar) using different biological matrices. (external link)
• Lene Sveen; Gerrit Timmerhaus; Aleksei Krasnov et al. (2019). Wound healing in post-smolt Atlantic salmon (Salmo salar L.). (external link)
• Roland Koedijk; Arild Folkvord; Atle Foss et al. (2010). The influence of first-feeding diet on the Atlantic cod Gadus morhua phenotype: survival, development and long-term consequences for growth. (external link)
• Tharmini Kalananthan; Ole Folkedal; Ana S. Gomes et al. (2023). Impact of long-term fasting on the stomach-hypothalamus appetite regulating genes in Atlantic salmon postsmolts. (external link)
• Bernat Morro; Richard Broughton; Pablo Balseiro Vigo et al. (2021). Endoplasmic reticulum stress as a key mechanism in stunted growth of seawater rainbow trout (Oncorhynchus mykiss). (external link)
• Inge Døskeland; Albert Imsland; Per Gunnar Fjelldal et al. (2016). The effect of low temperatures and photoperiods on growth and vertebra morphometry in Atlantic salmon. (external link)
• Albert Imsland; Klement Arild Våge; Sigurd O Handeland et al. (2011). Growth and osmoregulation in Atlantic salmon (Salmo salar ) smolts in response to different feeding frequencies and salinities. (external link)
• Floriana Lai; Ivar Rønnestad; Sergei Budaev et al. (2024). Freshwater history influences farmed Atlantic salmon (Salmo salar) performance in seawater. (external link)
• Sigurd O Handeland; Albert Imsland; Björn Thrandur Björnsson et al. (2013). Long-term effects of photoperiod, temperature and their interaction on growth, gill Na+, K+-ATPase activity, seawater tolerance and plasma growth-hormone levels in Atlantic salmon Salmo salar. (external link)
• Sara Calabrese; Albert K. D. Imsland; Tom Ole Nilsen et al. (2023). Water Flow Requirements of Post-smolt Atlantic Salmon (Salmo salar L.) Reared in Intensive Seawater Flow-through Systems: A Physiological Perspective. (external link)
• Albert Imsland; Sigurd O Handeland; Sigurd Olav Stefansson (2014). Photoperiod and temperature effects on growth and maturation of pre- and post-smolt Atlantic salmon. (external link)
• Bibbi Maria Kallay Hjelle; Albert Imsland; Pablo Balseiro Vigo et al. (2022). Acoustic Delicing of Atlantic Salmon (Salmo salar): Fish Welfare and Salmon Lice (Lepeophtheirus salmonis) Dynamics. (external link)
• Christian Karlsen; Elisabeth Ytteborg; Gerrit Timmerhaus et al. (2018). Atlantic salmon skin barrier functions gradually enhance after seawater transfer. (external link)
• Sigurd O Handeland; Albert Imsland; Sigurd Olav Stefansson (2008). The effect of temperature and fish size on growth, feed intake, food conversion efficiency and stomach evacuation rate of Atlantic salmon post-smolts. (external link)
• Pablo Balseiro Vigo; Øyvind Grøner Moe; Ingrid Gamlem et al. (2018). Comparison between Atlantic salmon Salmo salar post-smolts reared in open sea cages and in the Preline raceway semi-closed containment aquaculture system. (external link)
• Cairsty Grassie; Victoria Anne Braithwaite; Jonatan Nilsson et al. (2013). Aluminum exposure impacts brain plasticity and behavior in Atlantic salmon (Salmo salar). (external link)
• Jonas Ismael Silva Marrero; Floriana Lai; Sigurd Olav Handeland et al. (2025). Integration of Gill and Intestinal Osmoregulatory Functions to Assess the Smoltification Window in Atlantic Salmon. (external link)
• Are Nylund; Egil Karlsbakk; Stian Nylund et al. (2008). New clade of betanodaviruses detected in wild and farmed cod (Gadus morhua) in Norway. (external link)
• Tarald Kleppa Øvrebø; Pablo Balseiro Vigo; Albert Imsland et al. (2022). Investigation of growth performance of post-smolt Atlantic salmon (Salmo salar L.) in semi closed containment system: A big-scale benchmark study. (external link)
• Enrique Pino Martinez; Pablo Balseiro Vigo; Håkon Fagerbakke Kvittingen et al. (2024). Influence of salinity on rainbow trout (Oncorhynchus mykiss) smolt development and postsmolt performance. (external link)
• Albert Imsland; Bjørn Roth; Per Gunnar Fjelldal et al. (2017). The effect of continuous light at low temperatures on growth in Atlantic salmon reared in commercial size sea pens. (external link)
• Albert Imsland; Bjørn Roth; Inge Døskeland et al. (2019). Flesh quality of Atlantic salmon smolts reared at different temperatures and photoperiods. (external link)
• Sigurd O Handeland; Albert Imsland; Tom Ole Nilsen et al. (2014). Osmoregulation in Atlantic salmon Salmo salar smolts transferred to seawater at different temperatures. (external link)
• Gianmarco Del Vecchio; Floriana Lai; Ana S. Gomes et al. (2021). Effects of short term fasting on mRNA expression of ghrelin and the peptide transporters PepT1 and 2 in Atlantic salmon (Salmo salar). (external link)
• Sigurd O Handeland; Albert Imsland; Lars O.E. Ebbesson et al. (2013). Low light intensity can reduce Atlantic salmon smolt quality. (external link)
• Enrique Pino Martinez; Pablo Balseiro Vigo; Cindy Pinto Pedrosa et al. (2021). The effect of photoperiod manipulation on Atlantic salmon growth, smoltification and sexual maturation: A case study of a commercial RAS. (external link)
• Sergei Budaev; Magda L. Dumitru; Katja Enberg et al. (2024). Premises for a digital twin of the Atlantic salmon in its world: Agency, robustness, subjectivity and prediction. (external link)
• Arne Mikal Arnesen; Hilde Toften; Thorleifur Agustsson et al. (2003). Osmoregulation, feed intake, growth and growth hormone levels in 0+ Atlantic salmon (Salmo salar L.) transferred to seawater at different stages of smolt development. (external link)
• Sveinung Fivelstad; Kristin Kvamme; Sigurd Olav Handeland et al. (2015). Growth and physiological models for Atlantic salmon (Salmo salar L.) parr exposed to elevated carbon dioxide concentrations at high temperature. (external link)
• Ana S. Gomes; Pablo Balseiro Vigo; Mari Dybvad Iversen et al. (2025). Performance of Atlantic salmon reared under three different regimes of continuous aerobic exercise during the freshwater phase. (external link)
• Sigurd O Handeland; Albert Imsland; Björn Thrandur Björnsson et al. (2013). Physiology during smoltification in Atlantic salmon: effect of melatonin implants. (external link)
• Sigurd O Handeland; Albert Imsland; Lars O.E. Ebbesson et al. (2014). Osmoregulation and growth in offspring of wild Atlantic salmon at different temperatures. (external link)
• Bernat Morro; Pablo Balseiro Vigo; Amaya Albalat et al. (2019). Effects of different photoperiod regimes on the smoltification and seawater adaptation of seawater-farmed rainbow trout (Oncorhynchus mykiss): Insights from Na+, K+-ATPase activity and transcription of osmoregulation and growth regulation genes. (external link)
• Tharmini Kalananthan; Floriana Lai; Ana S. Gomes et al. (2020). The Melanocortin System in Atlantic Salmon (Salmo salar L.) and Its Role in Appetite Control. (external link)
• Wenda Cui; Eisuke Takahashi; Bernat Morro et al. (2022). Changes in circulating insulin-like growth factor-1 and its binding proteins in yearling rainbow trout during spring under natural and manipulated photoperiods and their relationships with gill Na+, K+-ATPase and body size. (external link)
• Bernat Morro; Pablo Balseiro Vigo; Amaya Albalat et al. (2019). Effects of temperature and photoperiod on rainbow trout (Oncorhynchus mykiss) smoltification and haematopoiesis. (external link)
• Sigurd O Handeland; E Wilkinson; B Sveinsbo et al. (2004). Temperature influence on the development and loss of seawater tolerance in two fast-growing strains of Atlantic salmon. (external link)
• Jarl Giske; Magda L. Dumitru; Katja Enberg et al. (2025). Premises for digital twins reporting on Atlantic salmon wellbeing. (external link)
• Bernat Morro; Mary K. Doherty; Pablo Balseiro Vigo et al. (2020). Plasma proteome profiling of freshwater and seawater life stages of rainbow trout (Oncorhynchus mykiss). (external link)
• Enrique Pino Martinez; Albert Kjartan Dagbjartarson Imsland; Anne-Camilla Diesen Hosfeld et al. (2023). Effect of Photoperiod and Transfer Time on Atlantic Salmon Smolt Quality and Growth in Freshwater and Seawater Aquaculture Systems. (external link)
• Enrique Pino Martinez; Kari Anne Kamlund; Pablo Balseiro Vigo et al. (2024). Elevated water temperature in combination with an increase in daylength stimulate a synchronized onset of spermatogenesis in male Atlantic salmon (Salmo salar, L.) postsmolts. (external link)
• Enrique Pino Martinez; Kristine Folde Staveland; Cindy Pinto Pedrosa et al. (2025). A decrease in water temperature from 15 °C to 8 °C before or after a “winter signal” photoperiod delays but does not arrest early sexual maturation in male Atlantic salmon Salmo salar L. postsmolts. (external link)
• Maren Skare; Sherry Stephanie Chan; Sigurd Olav Handeland et al. (2021). A comparative study on quality, shelf life and sensory attributes of Atlantic salmon slaughtered on board slaughter vessels against traditional land-based facilities.. (external link)
• Sara Calabrese; Tom Ole Nilsen; Jelena Kolarevic et al. (2017). Stocking density limits for post-smolt Atlantic salmon (Salmo salar L.) emphasis on production performance and welfare. (external link)
• Enrique Pino Martinez; Markus Førde Braanaas; Pablo Balseiro Vigo et al. (2022). Constant High Temperature Promotes Early Changes in Testis Development Associated with Sexual Maturation in Male Atlantic Salmon (Salmo salar L.) Post-Smolts. (external link)
• Lene Sveen; Gerrit Timmerhaus; Jacob Seilø Torgersen et al. (2016). Impact of fish density and specific water flow on skin properties in Atlantic salmon (Salmo salar L.) post-smolts. (external link)
• Enrique Pino Martinez; Pablo Balseiro Vigo; Sigurd Olav Stefansson et al. (2023). Interaction of temperature and feed ration on male postsmolt maturation of Atlantic salmon (Salmo salar L.). (external link)
• Bjørn Roth; Sten Johan Sverre Johansen; Jorma Suontama et al. (2005). Seasonal variation in flesh quality, comparison between large and small Atlantic salmon (Salmo salar) transferred into seawater as 0+ or 1+ smolts. (external link)
• Gunnar Steinn Gunnarsson; Steffen H. Blindheim; Egil Karlsbakk et al. (2017). Desmozoon lepeophtherii (microsporidian) infections and pancreas disease (PD) outbreaks in farmed Atlantic salmon (Salmo salar L.). (external link)
• Albert Imsland; Pablo Balseiro Vigo; Sigurd Olav Handeland et al. (2025). Follow-Up Study on Acoustic De-Licing of Atlantic Salmon (Salmo salar): Lepeophtheirus salmonis and Caligus elongatus Dynamics over Four Consecutive Production Cycles. (external link)
• Sigurd Olav Stefansson; Albert Imsland; Sigurd O Handeland (2009). Food-deprivation, compensatory growth and hydro-mineral balance in Atlantic salmon (Salmo salar) post-smolts in sea water. (external link)
• Camilla Diesen Hosfeld; Janicke Hammer; Sigurd O Handeland et al. (2009). Effects of fish density on growth and smoltification in intensive production of Atlantic salmon (Salmo salar L.). (external link)
• Enrique Pino Martinez; Sigurd Olav Handeland (2025). A Case Study Documenting Growth and Welfare of Atlantic Salmon Reared to Market Size in a Novel Commercial Hybrid Flow-Through System in Norway: A Comparison With Fish Performance in Traditional Sea Sites. (external link)
• Floriana Lai; Ivar Rønnestad; Thea Solberg Olsen et al. (2025). Adaptations to intermittent fasting in large sea caged Atlantic salmon (Salmo salar); effects on feeding, energy homeostasis, and growth. (external link)
• Enrique Pino Martinez; Kristine Folde Staveland; Mitchell Stewart Fleming et al. (2025). Body size of male Atlantic salmon (Salmo salar L.) at introduction of a 5-week LD12:12 winter signal influences their decision to mature early or smoltify. (external link)
• Tom Ole Nilsen; Lars O.E. Ebbesson; Sigurd O Handeland et al. (2013). Atlantic salmon (Salmo salar L.) smolts require more than two weeks to recover from acidic water and aluminium exposure. (external link)

Conference lecture

• Sara Calabrese; Sigurd O Handeland; Bjarte Sævareid et al. (2015). Semi-closed sea systems for postsmolt Atlantic salmon. Conference Abstract p. 123. (external link)
• Trine Ytrestøyl; Harald Rune Takle; Jelena Kolarevic et al. (2015). Atlantic salmon post-smolts in RAS: effects of salinity, exercise and timing of seawater transfer on performance, physiology and welfare. In Book of abstracts, p. 39. (external link)
• Bendik Fyhn Terjesen; W. Abbink; E Blom et al. (2015). Scaling of culture tanks and unit processes, relevant for Atlantic salmon post-smolt production in land-based systems. In Book of abstracts, p. 35. (external link)
• Sigurd O Handeland; Torbjørn Järvi; Anders Fernø (1993). Consequences of osmotic stress on the anti-predator behaviour and predator mortality of atlantic salmon (Salmo salar L.) postsmolts. (external link)
• Jelena Kolarevic; Britt Kristin Megård Reiten; Lene Sveen et al. (2016). Welfare and performance of Atlantic salmon post-smolts during exposure to mild chronic stress in recirculation aquaculture system (RAS).. (external link)
• Sveinung Fivelstad; Kristin Kvamme; Sigurd O Handeland et al. (2012). Karbondioksid og vekstreduksjon hos parr. (external link)
• Tom Ole Nilsen; H. Sundh; Eva Andersson et al. (2016). Osmoregulatory changes in sexually maturing Atlantic salmon post-smolt and adults. (external link)
• Enrique Pino Martinez; Asbjørn Dyrkorn Løland; Sigurd Olav Handeland (2024). COMPARING GROWTH AND WELFARE OF ATLANTIC SALMON POST-SMOLTS IN A NOVEL LAND-BASED COMMERCIAL HYBRID FLOW-THROUGH SYSTEM (HFS) VERSUS IN OPEN SEA PENS. (external link)
• Ana S. Gomes; Pablo Balseiro Vigo; Ivar Rønnestad et al. (2022). Effects of swimming exercise on perfomance of Atlantic salmon. (external link)
• Bendik Fyhn Terjesen; Sigurd O Handeland (2013). Postsmolt - ytelse og velferd i lukkede og semilukkede anlegg. (external link)
• Sigurd O Handeland; T. Järvi; Anders Fernø et al. (1991). Consequences of seawater acclimation on the survival of Atlantic salmon (Salmo Salar L.) smolts in the presence of cod (Gadus morhua L.). (external link)
• Christian Karlsen; Elisabeth Ytteborg; Gerrit Timmerhaus et al. (2018). Hudens utvikling og immunfunksjoner i postsmoltfasen. (external link)
• Sara Calabrese; Tom Ole Nilsen; Sveinung Fivelstad et al. (2014). The effects of increased CO2 on postsmolt Atlantic salmon Salmo salar growth, physiology and stress responsiveness. (external link)
• Jonas Ismael Silva Marrero; Ana S. Gomes; Pablo Balseiro et al. (2024). Assessing the physiological response of gills and intestine through smoltification in Atlantic salmon exposed to two different photoperiods. (external link)
• Trond Einar Isaksen; Egil Karlsbakk; Karl Fredrik Ottem et al. (2007). Screening av smittestoffer i vill stamfisk. (external link)
• Trine Ytrestøyl; Harald Rune Takle; Jelena Kolarevic et al. (2015). Atlantic salmon post-smolts in RAS: Effects of salinity, exercise and timing of seawater transfer on performance, physiology and welfare. (external link)
• Sigurd O Handeland; Sigurd Olav Stefansson; Lars O.E. Ebbesson et al. (2012). Økende temperaturer fremskynder smoltutvikling hos Vosso laks. Resultat fra prosjektet: "The smolt probe - novel tools for assessment of smolt quality and marine performance in Atlantic salmon (SmoltProbe)". (external link)
• Lene Sveen; Gerrit Timmerhaus; Jacob Seilø Torgersen et al. (2017). My skin don't lie.. (external link)
• Sigurd O Handeland (2017). Overview of CtrlAQUA research in post-smolt welfare, performance, and water quality in commercial scale, floating, semi-closed containment systems: CtrAQUA Preline, Neptune and Eco-Cage. (external link)
• Bendik Fyhn Terjesen; Trine Ytrestøyl; Jelena Kolarevic et al. (2013). Effects of salinity and exercise on Atlantic salmon postsmolts reared in land-based RAS. (external link)
• Sara Calabrese; Bjarte Sævareid; Olav Breck et al. (2016). Semi-closed sea systems for postsmolt Atlantic salmon. (Abstract ID 42049).. (external link)
• Kristin Kvamme; Camilla Diesen Hosfeld; Sigurd O Handeland et al. (2014). Effekter på helse sykdom og velferd ved postsmoltproduksjon i semi-lukkede anlegg – Døgnvariasjon i vannkvalitet i anlegg for postsmolt. (external link)
• Kristine Folde Staveland; Enrique Pino Martinez; Mitchell Stewart Fleming et al. (2023). INFLUENCE OF A TEMPERATURE DROP FROM 15 TO 8°C BEFORE OR AFTER WINTER SIGNAL ON EARLY SEXUAL MATURATION IN ATLANTIC SALMON Salmo salar L. POSTSMOLT. (external link)
• Jelena Kolarevic; Lene Sveen; Tom Ole Nilsen et al. (2016). Welfare and performance of Atlantic salmon post-smolts during exposure to mild chronic stress in closed-containment systems. In Aquaculture Europe 2016 Abstracts, pp 533-534. (external link)
• Sveinung Fivelstad; Camilla Diesen Hosfeld; sara calabrese et al. (2014). Effekter på helse, sykdom og velferd ved postsmoltproduksjon i semi-lukkede anlegg – del D: Grenseverdier for karbondioksid for postsmolt.. (external link)
• Christopher Good; S. Summerfelt; T. May et al. (2017). The effects of photoperiod on Atlantic salmon post-smolt in freshwater closed-containment systems. (external link)
• Pablo Balseiro Vigo; Erlend Nordvoll; Sara Calabrese et al. (2022). Performance of Salmo salar smolt in sulfate reduced nanofiltered water. (external link)
• Chris Good; S. Summerfelt; Brian Vinci et al. (2019). The Freshwater Institute & CtrlAQUA, Norway: A multi-year research collaboration on RAS-based Atlantic salmon production.. (external link)
• Sara Calabrese; Sigurd Olav Handeland; Bendik Fyhn Terjesen et al. (2014). Oppdrett av postsmolt laks i semilukkede system; Krav til fisketetthet og vannforbruk.. (external link)
• Tom Ole Nilsen; Eva Andersson; Stephen D. McCormick et al. (2016). Androgens up-regulate freshwater ion transporters in seawater acclimated Atlantic salmon (Salmo salar). (external link)

Media interview

• Pia Beate Pedersen; Sigurd O Handeland (2012). Unngå å stresse fisken ved transport. (external link)
• Sigurd Olav Handeland; Jan-Egil Dyvik (2011). Dale Klekkeri blir feltlab. (external link)
• Lars O.E. Ebbesson; Sigurd O Handeland (2015). Tror lukkede anlegg vil være til hjelp for oppdretterne i framtida. (external link)
• Sigurd O Handeland; Tom Ole Nilsen (2015). Nytt analyseverktøy skal sikre riktig tidspunkt for smoltutsett. (external link)
• Therese Soltveit; Sigurd O Handeland (2016). – Dypere vann gir bedre vekstrate for postsmolt. (external link)
• Lars Kvamme; Sigurd O Handeland (2014). Kan få mye å si for lusesituasjonen. (external link)
• Harald Berglihn; Sigurd Olav Handeland (2014). Minimalt svinn i lukket forskningsanlegg. (external link)
• Linn Therese Skår Hosteland; Sigurd O Handeland; Tom Ole Nilsen (2015). Satser på histologi, rognkjeks og ny smoltanalyse. (external link)
• Eivor Jerpåsen; Sigurd O Handeland (2016). Mener de har løsningen på laksenæringens milliardproblem. (external link)
• Øyvind Sjøthun Røen; Sigurd Olav Handeland (2015). Maner til innsats etter AGD-påvisning. (external link)
• Tore Stensvold; Sigurd O Handeland (2014). Nofima skal lede nytt forskningssenter for fiskeoppdrett. (external link)
• Sigurd Olav Handeland; Magnus Petersen (2016). - Kontinuerlig lys gir 30 % bedre tilvekst på lave temperaturer. (external link)
• Sigurd O Handeland (2013). Verdas største merd skal revolusjonera fiskeindustrien. (external link)
• Sigurd O Handeland; Lars O.E. Ebbesson (2016). Lovende resultater for produksjon av laks i lukkede anlegg. (external link)
• Linn Therese Skår Hosteland; Sigurd O Handeland; Hans Torstein Kleivdal (2017). 55 millioner til prosjekt for havbruksvekst i Hordaland. (external link)

Lecture

• Sigurd Olav Handeland (2024). Velferd i norsk havbruk. (external link)
• Sigurd Olav Handeland (2021). Potensielle utfordringer ved produksjon av laksefisk i RAS. (external link)
• Sigurd Olav Handeland (2021). Potensielle utfordringer ved produksjon av laksefisk i RAS.. (external link)
• Sigurd Olav Handeland (2021). Produksjon av robust postsmolt i et fremtidig perspektiv.. (external link)
• Sigurd Olav Handeland (2021). Tidlig modning hos postsmolt i RAS. (external link)
• Sigurd Olav Handeland (2024). Tidlig modning hos postsmolt av laks. (external link)
• Sigurd Olav Handeland (2024). Tidlig modning hos postsmolt. (external link)
• Sigurd Olav Handeland (2023). Postsmolt modning hos laks. (external link)
• Sigurd Olav Handeland (2013). "Effekt av ulike typer lys på fisk i settefiskanlegg". (external link)
• Sigurd Olav Handeland (2023). Postsmolt modning i RAS. (external link)
• Sigurd Olav Handeland (2024). Intensiv produksjon av smolt i RAS. (external link)
• Sigurd Olav Handeland (2024). Bærekraftig havbruk. (external link)
• Sigurd Olav Handeland (2024). Fiskevelferd i bærekraftig havbruk. (external link)
• Sigurd Olav Handeland (2024). Velferd i bærekraftig havbruk. (external link)
• Sigurd Olav Handeland (2024). Intensiv produksjon av laks. (external link)
• Sigurd Olav Handeland (2022). Tidlig kjønnsmodning hos laksefisk. (external link)
• Sigurd Olav Handeland (2024). Intensiv produksjon av postsmolt. (external link)
• Sigurd Olav Handeland (2013). Foredrag på FHF-samling Verdikjede havbruk, den 21.-22. oktober 2013: "Produksjon av laks i lukkede anlegg". (external link)
• Floriana Lai; Sigurd Olav Handeland; Pablo Balseiro Vigo et al. (2022). Sesongvariasjon av fôringsatferd, appetitt, vekst og fôrutnyttelse gjennom en produksjonssyklus av laks- tidlige livsstadier påvirker fisken helt frem til slakt. (external link)
• Floriana Lai; Ivar Rønnestad; Sergei Budaev et al. (2023). Seawater growth performance of Atlantic salmon (Salmo salar) smolts produced in a flow-through or recirculating aquaculture system.. (external link)
• Tom Ole Nilsen; Bendik Fyhn Terjesen; Jelena Kolarevic et al. (2018). Effects of high environmental CO2 levels on Atlantic salmon post-smolts. (external link)
• Bernat Morro; Mary K. Doherty; Pablo Balseiro Vigo et al. (2018). Changes in plasma proteome and peptidome during smoltification and seawater adaptation of steelhead trout (Oncorhynchus mykiss). (external link)
• Tharmini Kalananthan; Floriana Lai; Sigurd Olav Handeland et al. (2021). Appetittstyrt fôring av Atlantisk laks. (external link)
• Bernat Morro; Mary K. Doherty; Pablo Balseiro Vigo et al. (2018). Mass Spectrometry-Based Plasma Proteomics for Steelhead Trout (Oncorrhynchus mykiss) Aquaculture. (external link)
• Bernat Morro; Mary K. Doherty; Pablo Balseiro Vigo et al. (2018). Plasma and liver proteome of an unwanted phenotype of seawater-transferred rainbow trout (Oncorhynchus mykiss). (external link)
• Bernat Morro; Mary K. Doherty; Pablo Balseiro Vigo et al. (2018). Plasma proteome during smoltification and seawater adaptation of steelhead trout (Oncorhynchus mykiss). (external link)
• Sigurd O Handeland (2010). Smoltkvalitet - Er smoltkvaliteten i negativ utvikling. (external link)
• Lene Sveen; Elisabeth Ytteborg; Gerrit Timmerhaus et al. (2016). Effect of stress on wound healing and gene regulation in skin of posts-smolts (Salmo salar L.) in closed-containment aquaculture systems (in Norwegian). In Abstracts RCN Program conference Mariculture 2016. (external link)
• Astrid Buran Holan; Brian Vinci; Sigurd O Handeland et al. (2016). Evaluation of available water treatment technologies for semi-closed aquaculture systems (in Norwegian). In Abstracts RCN Program conference Mariculture 2016. (external link)
• Pablo Balseiro Vigo; Øyvind Moe; Ingrid Gamlem et al. (2017). Muscle development and heart condition in Atlantic salmon (Salmo salar) post-­smolts reared in Preline S-CCS. (external link)
• Floriana Lai; Sigurd Olav Handeland; Pablo Balseiro Vigo et al. (2022). Seasonal variation of feeding behaviour, appetite, growth and feed utilization through a salmon production cycle.. (external link)
• Enrique Pino Martinez; Pablo Balseiro Vigo; Markus Braanaas et al. (2019). INFLUENCE OF WATER TEMPERATURE AND FEEDING REGIME ON THE INCIDENCE OF EARLY SEXUAL MATURATION IN ATLANTIC SALMON (Salmo salar L.) POSTSMOLTS DURING THE FRESHWATER STAGE. (external link)
• Sigurd Olav Handeland (2023). Fra villaks til husdyr. (external link)
• Sigurd Olav Handeland (2022). Postsmoltens behov i lukkede systemer for å sikre en optimal tilværelse i sjø. (external link)
• Sigurd Olav Handeland (2023). RASande god postsmolt. (external link)
• Sigurd Olav Handeland (2023). Optimalisering av nye oppdrettsprotokoller (Hybrid Flow through System) med fokus på tilvekst, velferd og økonomisk prestasjon. (external link)
• Sigurd Olav Handeland (2021). Produksjon av robust postsmolt. (external link)

Doctoral thesis (PhD)

• Sigurd O Handeland (1998). Osmoregulation and growth in Atlantic salmon smolts during the early seawater phase. (external link)
• Sigurd Olav Handeland; Bernat Morro (2019). Development of a full season production protocol for rainbow trout (Oncorhynchus mykiss). (external link)
• Sigurd Olav Handeland; Sara Calabrese (2017). Environmental and biological requirements of post-smolt Atlantic salmon (Salmo salar L.) in closed-containment aquaculture systems. (external link)
• Sigurd Olav Handeland; Enrique Pino Martínez (2021). Influence of water temperature, photoperiod and feeding regime on early sexual maturation of Atlantic salmon (Salmo salar L.) postsmolts during the freshwater phase. (external link)
• Sigurd Olav Handeland; Bøe Beate Louise; Jordan Sivertsen Petter (2021). Tilvekst og fiskevelferd ved bruk av sulfatredusert brakkvann i RAS-anlegg. (external link)
• Sigurd Olav Handeland; Roland Koedjijk (2009). Phenotypic plasticity influenced by diet during early development : A comparative study on Atlantic cod. (external link)
• Sigurd Olav Handeland; Ole Bergheim; Inger Lund (2009). Transportforsøk på torsk- Gadus morhua. (external link)
• Sigurd Olav Handeland; Anne Lise Fløtre; Detelina Manolova Hristova et al. (2010). Grenseverdi for karbondioksid for torskeyngel (Gadus morhua). (external link)
• Sigurd Olav Handeland; Camilla D. Hosfeld (2010). Physiological effects of environmental hyperoxia on Atlantic salmon (Salmo salar L.) and European sea bass (Dicentrarchus labrax L.).. (external link)
• Sigurd Olav Handeland; Lene Sveen (2018). Aquaculture relevant stressors, impacts on skin and wound healing in Atlantic salmon (Salmo salar L.) postsmolts. (external link)
• Sigurd Olav Handeland; Caroline Ulvesæter; Nathalie Reiertsen (2020). Valg av overflatebehandling for landbaserte anlegg. Hovedoppgave Havteknologi. (external link)

Professional article

• Sara Calabrese; Sigurd O Handeland; Bendik Fyhn Terjesen et al. (2014). Krav til fisketetthet og vannforbruk. (external link)
• Sigurd O Handeland; Henning Andre Urke; Torstein Kristensen et al. (2008). Påvirker vannkvalitet og produksjonsintensitet risiko for utvikling av vintersår hos postsmolt - Resultater fra en storskala feltstudie. (external link)
• Floriana Lai; Ivar Rønnestad; Pablo Balseiro Vigo et al. (2022). Dagens intensive produksjon av laks er ikke tilpasset laksens biologi. (external link)

Academic book chapter

• Sigurd Stefansson; Grete Bæverfjord; Roderick Nigel Finn et al. (2006). Vannkvalitet - laksefisk. (external link)
• Tom Ole Nilsen; Bendik Fyhn Terjesen; Jelena Kolarevic et al. (2018). Effects of high environmental CO2 levels on Atlantic salmon post-smolts. (external link)

Conference poster

• Helene Mikkelsen; Hilde Toften; Børge Damsgård et al. (2007). Increased risk of winter ulcer in intensively produced Atlantic salmon juveniles exposed to seawater. (external link)
• Floriana Lai; Tharmini Kalananthan; G. M. Berg et al. (2019). Brain Distribution of Key Neuropeptides Involved in Appetite Control in Atlantic Salmon, Salmo Salar (L.).. (external link)
• Floriana Lai; Ida Hundven; Pablo Balseiro Vigo et al. (2023). Feed intake, growth and gut transit in Atlantic salmon at different temperatures- involvement of control pathways for digestion and appetite.. (external link)
• Bjørn Olav Rosseland; Trond Rosten; Brit Salbu et al. (2006). Dyrehelse og dyrevelferd i en intensiv produksjon.Vannkvalitetskriterier og effekter på fisk som kan brukes til å "måle" eller registrere velferd under ulike produksjonsforhold fra befruktning til sjøsetting og transport og opphold i sjø. (external link)
• Åsa Maria Olofsdotter Espmark; Jelena Kolarevic; Lill-Heidi Johansen et al. (2019). CtrlAQUA SFI – Contribution to Future Aquaculture. (external link)
• Sara Calabrese; Tom Ole Nilsen; Sigurd O Handeland et al. (2016). Farming intensification - impact on mental and physiological robustness of post-smolt Atlantic salmon (Salmo salar). (external link)
• Mitchell Stewart Fleming; Enrique Pino Martinez; Trine Hansen et al. (2021). Effect of water temperature and photoperiod on brain and pituitary maturation hormones during freshwater production stage of Atlantic salmon (Salmo salar L.). (external link)
• Enrique Pino Martinez; Kristine Folde Staveland; Mitchell Stewart Fleming et al. (2023). BODY SIZE OF MALE ATLANTIC SALMON INTERACTS WITH CHANGES IN PHOTOPERIOD UNDER AQUACULTURE CONDITIONS TO INFLUENCE THE DECISION TO MATURE OR SMOLTIFY. (external link)
• Patrik Tang; Tom Ole Nilsen; Naouel Gharbi et al. (2019). How Does Temperature Affect Primary Barrier Functions in Atlantic Salmon (Salmo Salar L.) Post-Smolts?. (external link)
• Patrik Tang; Tom Ole Nilsen; Henrik Sundh et al. (2019). The effect of temperature on stress disturbance and skin barrier functions in Atlantic salmon (Salmo salar L. ) post- smolts. (external link)

Master’s thesis

• Sigurd Olav Handeland; Tarald Kleppa Øvrebø (2020). Growth performance and welfare of post-smolt (Salmo salar L.) reared in semi closed containment systems (S-CCS) – a comparative study. (external link)
• Sigurd Olav Handeland; Lisa Gaassand Hagevik (2023). The effect of aerobic training on immune response in post-smolt Atlantic salmon (Salmo salar) when infected by salmon lice (Lepeophtheirus salmonis) in small-scale close containment system. (external link)
• Sigurd Olav Handeland; Bibbi Maria Kállay Hjelle (2021). Field study on Atlantic salmon (Salmo salar) acoustic delicing: Fish welfare and salmon lice dynamics. (external link)
• Sigurd Olav Handeland; Trine Beate Tangeraas Hansen (2022). The effects of water temperature and photoperiod: Investigating the cause of early activation of the BPG axis in male Atlantic salmon (Salmo salar) during the freshwater stages of development. (external link)
• Sigurd Olav Handeland; Fagerbakke Kvittingen Håkon (2019). Salinity influence on smolt development and post-smolt performance in Rainbow trout (Oncorhynchus mykiss). (external link)
• Sigurd Olav Handeland; Norvoll Erlend (2022). Performance of salmon post-smolt reared in ion modified brackish water in recirculating aquaculture systems. (external link)
• Sigurd Olav Handeland; Gunnar Merckoll Berg (2018). Neuroendocrine factors involved in appetite control and feed intake in salmon.. (external link)
• Sigurd Olav Handeland; Henrik Rye Jakobsen (2012). The effect of salinity on growth, appetite and feed conversion efficiency of Atlantic salmon post-smolts; expression of selected ion and nutrient uptake transporters in the anterior intestine. (external link)
• Jannicke Hammer; Sigurd Stefansson; Sigurd O Handeland (2007). Effects of stocking density on smoltification and growth of Atlantic salmon (Salmo salar L.). (external link)
• Sigurd Olav Handeland; Ronja Rogerdatter Athammer (2023). Tolerance limits to fluctuating water currents in Atlantic salmon (Salmo salar): A novel method to simulate the impact of ocean waves on salmon welfare in the laboratory. (external link)
• Sigurd Olav Handeland; Mari Dybvad Iversen (2023). Growth of Atlantic salmon (Salmo salar) under different regimes of continuous aerobic exercise. (external link)
• Sigurd Olav Handeland; Eidskrem Gullaksen Marianne (2023). Effect of continuous aerobic exercise on the heart of Atlantic salmon. (external link)
• Sigurd Olav Handeland; Øyvind Grøner Moe (2017). Muscle development and heart condition in Atlantic salmon (Salmo salar) post-smolts reared in Preline S-CCS.. (external link)
• Sigurd Olav Handeland; Ingrid Gamlem (2017). Documentation of growth and insulin-like growth factor I in Atlantic salmon (Salmo salar) post-smolts reared in large scale semi-closed and open systems. (external link)
• Sigurd Olav Handeland; Egor A. Gaidukov (2017). Effects of rapid temperature changes on Atlantic salmon, Salmo salar, post-smolts. (external link)
• Sigurd Olav Handeland; Tilde Sørstrand Haugen (2020). The effect of winter-signal photoperiod manipulations on smoltification of Atlantic salmon (Salmo salar) reared in commercial Recirculating Aquaculture Systems (RAS). (external link)
• Sigurd Olav Handeland; Kari Anne Kamlund (2021). The effect of temperature and photoperiod on early sexual maturation in male Atlantic salmon (Salmo salar L.) during the freshwater phase. (external link)
• Sigurd Olav Handeland; Inge Døskeland (2015). The effect of low temperatures and photoperiods on growth in Atlantic salmon (Salmo salar). (external link)
• Sigurd Olav Handeland; Skare Maren (2021). A comparative study on quality, shelf life and sensory attributes of Atlantic salmon slaughtered onboard slaughter vessel against traditional land-based facilities. (external link)
• Sigurd Olav Handeland; Markus Førde Braanaas (2021). Influence of temperature and feeding on early sexual maturation commitment in male Atlantic salmon (Salmo salar, L.) during the freshwater stage. (external link)
• Sigurd Olav Handeland; Jannicke Hammer (2007). Effects of stocking density on smoltification and growth of Atlantic salmon (Salmo salar L.).. (external link)
• Sigurd Olav Handeland; Kaja Andersen (2009). Simulated and large- scale transportation of juvenile Atlantic cod (Gadus morhua), with or without aeration and at different densities,. (external link)

Conference abstract

• Floriana Lai; Tharmini Kalananthan; G. M. Berg et al. (2019). Brain Distribution of Key Neuropeptides Involved in Appetite Control in Atlantic Salmon, Salmo Salar (L.).. (external link)

Research report

• Sigurd O Handeland; T. Järvi; Anders Fernø et al. (1991). Consequences of seawater acclimation on the survival of Atlantic salmon (Salmo Salar L.) smolts in the presence of cod (Gadus morhua L.). (external link)
• Wenche Kristin Farstad; Jon Martin Arnemo; Bjarne Olai Braastad et al. (2006). Dyrevelferdsmessig risikovurdering av produksjon av 0-årssmolt. Uttalelse fra Faggruppe for Dyrehelse og Dyrevelferd (Dyrevern) i Vitenskapskomiteen for mattrygghet. (external link)
• Ragnar Tveterås; Bård Misund; Sigurd Olav Handeland (2022). Høringssvar – etablering av en ny ordning for tildeling av miljøteknologitillatelser. (external link)
• Sigurd O Handeland; Sveinung Fivelstad; Bendik Fyhn Terjesen (2015). Effects on health, disease, and welfare of postsmolt production in semi-closed containment systems – Part A: Fish density tolerance and specific water use. (external link)
• Sigurd O Handeland; Sveinung Fivelstad; Bendik Fyhn Terjesen (2014). Effekter på helse, sykdom og velferd ved postsmoltproduksjon i semi-lukkede anlegg-FHF 900816. Sluttrapport.. (external link)
• Sveinung Fivelstad; Camilla Diesen Hosfeld; Sigurd O Handeland et al. (2015). Sluttrapport på postsmolt D: Grenseverdier for karbondioksid for postsmolt. (external link)
• Sigurd O Handeland; E. Wilkinsson; Sigurd Stefansson (2001). Smolting and de-smolting at different temperatures in two strains of Atlantic salmon (Salmo salar L.). (external link)
• Sigurd O Handeland (2001). Effekt av utsettingstid og tilleggsbelysning på smoltkvalitet og vekst i ferskvann og sjøvann; hvilke egenskaper er best egnet til å predikere utsett av smolt?. (external link)

Popular science article

• Tom Ole Nilsen; Juliane Lukas; Ragnhild Valen et al. (2015). Utvikling av osmoreguleringssystemet hos torskelarver startfôret på rotatorier eller naturlig zooplankton. (external link)

Letter to the editor

• Bendik Fyhn Terjesen; Sigurd O Handeland (2014). Fremtidens oppdrettslaks er både i lukkede og åpne anlegg. (external link)

Popular science book chapter

• S. O Stefansson; G Bæverfjord; Nigel Finn et al. (2007). Water quality-salmonids. (external link)
• S Stefansson; W Bjerknes; P Bjørn et al. (2007). Physiological quality of eggs, fingerlings and smolt. (external link)

Media feature article

• Bendik Fyhn Terjesen; Sigurd O Handeland (2014). Skjerm laksen mot lus. (external link)
• Hans Vanhauwaert Bjelland; Ragnar Tveterås; Bård Misund et al. (2024). Hvordan reduserer vi dødeligheten i havbruk?. (external link)

See a complete overview of publications in NVA.

Dr. scient. Institutt for fiskeri og marinbiologi, Universitetet i Bergen (1998)

Cand. Scient. Inst. for fiskeri og marinbiologi, Universitetet i Bergen (1993)